Heloniopsis (A.Gray) N.Tanaka

Tanaka, Noriyuki, 2019, Taxonomy, evolution and phylogeography of the genus Helonias (Melanthiaceae) revisited, Phytotaxa 390 (1), pp. 448-450 : 448-450

Sect. 2. Heloniopsis (A.Gray) N.Tanaka , comb. & stat. nov.

Type:― Helonias orientalis (Thunb.) N.Tanaka (≡ Scilla orientalis Thunb. = Heloniopsis pauciflora A.Gray ).

≡ Heloniopsis Gray (1859: 416) , excl. charact. seminum, nom. cons. Type:― H. pauciflora A.Gray.

= Hexonix Rafinesque (1837 , as ‘1836’: 13), nom. rejic. vs. Heloniopsis Gray (1859) (nom. cons.). ≡ Kozola Rafinesque (1837 , as ‘1836’: 25), nom. rejic. vs Heloniopsis Gray (1859) (nom. cons.). Basionym of types:― Scilla japonica Thunb.

= Sugerokia Miquel (1866: 24 ; 1867a: 88; 1867b: 144). Type:― Sugerokia japonica Miq.

= Ypsilandra Franchet (1887 – 1888: 93, t. 17). Type:― Y. thibetica Franch.

Japanese name:―Shôjôbakama setsu.

Species included:―11; Helonias alpina , H. breviscapa , H. jinpingensis , H. kawanoi , H. koreana , H. leucantha , H. orientalis , H. parviflora , H. thibetica , H. umbellata , H. yunnanensis .

Description:―Leaves hysteranthous, sometimes proteranthous. Inflorescences racemose or (sub)umbellate. Flowers 1–many, adichogamous or protogynous. Tepals adaxially nearly flat or canaliculate toward base, distinct, or basally connate to adjoining tepals and adnate to opposing filaments. Nectary at base of tepal nearly flat, slightly or moderately concave or subtubular. Filaments distinct or adnate proximally to opposing tepals; inner filaments free or adnate basally to base of ovary. Anthers basifixed or (sub)dorsi-basifixed, unilocular or appearing bilocular. Style (fused columnar portion) 0.2–22 mm. Stigma trisected, trilobed, discoid or (sub)capitate. Seeds linear-fusiform.

Distribution:― Bhutan (NE), China (SW), India (NE), Korea, Japan, Myanmar (N), Nepal (Central, E), Russia (E: S Sakhalin), Taiwan, Vietnam (N) (Figs. 15, 29, 31-2–4, 32-2–4).

Remarks:― Helonia sect. Heloniopsis consists of two subsections, H. subsect. Ypsilandra and H. subsect. Heloniopsis . They are compared in Table 5, and a key to separate them is below.

Identification key to the subsections of H. sect. Heloniopsis

1.