Plusioglyphiulus phra, Golovatch & Geoffroy & Mauriès & Vandenspiegel, 2011

Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & Vandenspiegel, Didier, 2011, The millipede genus Plusioglyphiulus Silvestri, 1923 in Thailand (Diplopoda, Spirostreptida, Cambalopsidae) 2940, Zootaxa 2940 (1), pp. 1-63 : 36-37

Plusioglyphiulus phra sp. n. Figs 32−35.

Material examined: Holotype male ( MNHN GA 069 ), Thailand, Huai Hin Prov. , Ban Hua Lap Distr., Cave Tham Mai Lap Lae, hand collection, 28 July 1987, leg. P. Leclerc (THA-HH-04) . Paratypes: 1 male ( MNHN GA 069 ) , 1 male ( SEM), same locality, together with holotype .

Name: To emphasize “phra”, in Thai a honorific prefix meaning anything associated with royalty, religion, mythology or celestial objects, here a noun in apposition.

Diagnosis: Differs from congeners in the oligotrichous gnathochilarium, coupled with the particular carinotaxy patterns, and the special shapes and armature of both gonopod pairs (see also Key below).

Description: Length 26–28 mm, width 1.1–1.2 mm, collum and segments in posterior third of body being equally broadest. Coloration grey-yellow, in places mottled marbled (red-) brown; head, antennae (except for antennomeres 5 and 6), collum, venter and legs light yellowish; metatergal crests and, especially, poriferous tubercles usually infuscate, dark brown.

Body with 57–58p+3–1a+T. Length of holotype about 26 mm, width 1.1 mm, with 57p+3a+T.

All characters as in P. panhai sp. n. ( Figs 32E, F, 33B, C, F, 35A, E), except as follows. Ocellaria small, brownish, obviously partly faded towards peripheries, ovoid in shape, with about 12–14 visible ocelli arranged in 4–5 longitudinal rows. Gnathochilarium ( Fig. 35B) rather oligotrichous, with a separate promentum (n=2).

Postcollum constriction very evident, due to especially enlarged collum and segment 2 ( Figs 32A, D). Carinotaxy formula of collum: //(t)+/(t)/+1p/t+2p/t+(t)/(t)/(t)+3p/(t)/t+pp/t/t+/ma/m ( Figs 32A, D, 33D). Carinotaxy of metaterga 2–4, 7/7+m/m+7/7; usual formula of all following metaterga until about caudal third of body, 3/3+I/i+3/ 3/3+m/m/m, thereafter, 4/4+I/i+3/3/3+m/m/m ( Figs 32A–C, 33E); all crests and tubercles, including poriferous cones, rather low. Midbody segments ovoid in cross-section, slightly compressed laterally ( Fig. 33E).

Male legs 1 ( Fig. 35C) with a usual strong central hook curved forward, appendages larger, sac-shaped, with a few setae mesally near base, 1-segmented. Male legs 2 strongly enlarged ( Figs 33A, 35D).

Anterior gonopods rather complex, with a paramedian pair of anterior coxosternal processes (cxp1), relatively high, rather slender, largely contiguous, slightly curved, setose apically and carrying a subterminal tooth on front face; caudal pair of coxosternal processes (cxp2) higher, twisted and pointed; telopodites (te) club-shaped, setose apically, considerably shorter than cxp2, attached to coxal region caudolaterally, probably capable of movement ( Figs 34A–D, 35F). Posterior gonopods considerably shorter, rather simple, coxites well separated from sternum, fused only basally, sac-shaped, membranous, each surmounted on frontal face by an evident fovea devoid of nearby flagella, but supplied with a small tooth medio-apically; telopodites missing ( Figs 34E, F, 35G).