Plusioglyphiulus jaydee, Golovatch & Geoffroy & Mauriès & Vandenspiegel, 2011

Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & Vandenspiegel, Didier, 2011, The millipede genus Plusioglyphiulus Silvestri, 1923 in Thailand (Diplopoda, Spirostreptida, Cambalopsidae) 2940, Zootaxa 2940 (1), pp. 1-63 : 40-43

Plusioglyphiulus jaydee sp. n. Figs 36−38.

Material examined: Holotype male ( MNHN GA 070 ), Thailand, Kanchanaburi Prov., Sai Yok Distr., Ban Nam Tok, Cave Tham Rawa , hand collection, 16 June 1986, leg. P. Leclerc (THA-KAN-039) . Paratype: 1 male ( MNHN GA 070 ) , 1 female ( SEM), same locality, together with holotype .

Name: To emphasize “jaydee”, in Thai meaning “stupa, pagoda or chedi”, a noun in apposition to reflect the normally strongly bulging collum in Plusioglyphiulus .

Diagnosis: Differs from congeners in the oligotrichous gnathochilarium showing only a weak border between the pro- and eumentum, coupled with the presence of 2+2 dorsal tubercles on the epiproct, the particular carinotaxy patterns, and the special shapes and armature of both gonopod pairs (see also Key below).

Description: Collum and segments in posterior third of body equally broadest. Coloration uniformly yellowbrown; head, antennae, collum, venter and legs light yellowish; metatergal crests and, especially, poriferous tubercles only slightly infuscate, brown.

Length of holotype about 22 mm, width 1.1 mm, with 52p+3a+T. Length of male paratype about 21 mm, width 1.1 mm, with 47p+3a+T.

All characters as in P. panhai sp. n. ( Figs 36D, F, 37B, C, F, 38D), except as follows. Ocellaria small, brownish, obviously partly faded at peripheries, ovoid in shape, with about 12–14 visible ocelli arranged in 4–5 longitudinal rows. Gnathochilarium ( Fig. 38A) rather oligotrichous, with an indistinctly separated promentum (n=1).

Postcollum constriction very evident, due to especially enlarged collum and segment 2 ( Figs 36A, C, 37A). Carinotaxy formula of collum: /(t)/(t)+/(t)/+1p/t+/(t)/(t)+2p/t+3p/t/t+/t/t+pp/t/t+/ma/m ( Figs 36A, C). Carinotaxy of metaterga 2–4, 7/7+m/m+7/7; usual formula of all following metaterga, 3/3+I/i+3/3/3+m/m/m ( Figs 36A, B, 37E); all crests and tubercles, including poriferous cones, rather low. Midbody segments ovoid in cross-section, slightly compressed laterally ( Fig. 37E). Epiproct dorsally with 2+2 tubercles arranged in a transverse row ( Figs 36E, 37D).

Male legs 1 ( Fig. 38B) with a usual strong central hook curved forward, appendages rather small, sac-shaped, poorly setose, 2-segmented. Male legs 2 strongly enlarged, penes bare ( Fig. 38C).

Anterior gonopods rather simple, with a paramedian pair of anterior coxosternal processes (cxp1), relatively low, rather slender, contiguous, laterally sparsely setose, apically narrowly rounded; caudal pair of coxosternal processes (cxp2) higher, twisted, unciform, pointed; telopodites (te) club-shaped, setose apically, considerably shorter than cxp2, attached to coxal region caudolaterally, probably capable of movement ( Fig. 38E). Posterior gonopods considerably shorter, rather simple, coxites well separated from sternum, fused only basally, sac-shaped, membranous, each surmounted by a large fovea supplied with a short, but evident flagellum on top; telopodites missing ( Figs 38F, G).

Remarks: The unusually feeble border-line between the pro- and eumentum observed in the single dissected sample so vividly reminds us of several such cases in Glyphiulus species that it forces us to use this trait with caution as a species-specific character in Plusioglyphiulus as well. It has thus been omitted from the Key below.