Niladmirara Fedotova, 2018

Fedotova, Z. A., 2018, New taxa of the gall midges of the tribe Asynaptini (Diptera: Cecidomyiidae, Porricondylinae) from the Russian Far East, Far Eastern Entomologist 362, pp. 1-12 : 3

Genus Niladmirara Fedotova , gen. n.

Type species. Niladmirara metula Fedotova , sp. n., designated here.

DESCRIPTION. MALE. Body yellow, pale brown or brown with dark notum.

Length of wing 2.18–3.20 mm. Eye bridge 5-14 ommatidia-broad. Antennae 2+15-

17 or 2+21-22 segments. Face yellow. Mouth parts short, segments of palpi very narrow. Labrum pointed, labellum round. Scape, pedicel and flagellomeres uniformly sclrotized. Middle flagellomeres with subcylindric or barrel-shaped basal enlargement, neck 1.2–1.8 times as long as basal enlargement. Basal enlargement with simple ring sensorial file, large horse-shaped sockets in one row. Necks of middle flagellomeres without subapical swollen, basal enlargement with transparent almost invisible minute microtrichiae, or microtrichiae absence ( Niladmirara mucronata ).

Ultimate segment small, sometimes fused with penultimate one. Palpi 4-segmented,

much longer than length of head ( Fig. 9 ).

Thorax slightly sclerotized, uniformly yellow-brown. Notum brown, darker than thorax, with long lateral setae. Scutellum and last part of thorax uniformly sclerotized.

Hind legs shorter than fore and middlelegs. All femur slightly longer than tibiae.

Claws in all tarsi with dent, empodium as long as claws, slightly shorter or longer than ones. All veins distinct, R 5 strongly curved near apex, cell between veins C and

R 5 very naerrow. CuA 1 and CuA 2 well developed. Length of wing 3.2–3.3 times as long as width. Apex of wing slightly narrowed.

Abdomen tergites and sternites whole, with distal row of setae. Genitalia wide transversal or almost round. Gonocoxites very wide, thick, with truncated apex,

strongly enlarged distally; completely fused ventrally, almost without excision or with semiround excision. Gonostylus very wide, strongly enlarged dorsally, densely covered by swollen conical pores with very long setae ( Figs 20, 23 ), with small light claw that pointed or round apically. Gonostylus ventrally near claw with excision and short setae ( Figs 20, 24 ). Tegmen with shallow excision, slightly narrower than gonocoxites, truncated apically. IX tergite cordiform, slightly narrower than tegmen.

X tergite short, with truncated or round lobes and triangular excision, densely covered by microtrichiae ( Figs 21, 22 ). Ventral side of gonocoxites with medial slightly sclerotized aedeagal capsule. Aedeagus cylindrical, almost transversal or invisible. Parameres forked, with two pairs overlapping very long and thin projections; roots of parameres usually straight and thin. Genital rod long and thin,

almost parallelsided, right truncated apically ( Figs 21, 22 ). Basal part of genital rod sclerotized and enlarged on the end, somethimes far prolonging behind basal margin of genitalia ( Figs 1 , 10 ). This enlarged part slightly curved around rod.

Apical part of genital rod is surrounded by aedeagal capsule.

FEMALE (known for Niladmirara mucronata only). Antennae with 2+21 to

2+26 segmented; segments with short neck; sensoriae forming 2 rings; ovipositor curved upwards; terminal segment of lamella with long bristles.

SPECIES INCLUDED. The genus consists of three species distributed in the

Russian Far East: Niladmirara mucronata (Mamaev et Zaitzev, 1998) , comb. n.

Figures

Figs 1–9. Niladmirara metula sp. n., male holotype. 1 – genitalia; 2 – F5; 3 – F9; 4 – F13-F16 of left antenna; 5 – F14-F15 of right antenna; 6–7 – tarsal claw; 8 – wing; 9 – head. Scale bars = 0.1 mm.

Figs 17–24. Niladmirara spp., male holotypes: 17–21 N. devergentia sp. n.; 22–23 N.

Figs 10–16. Niladmirara devergentia sp. n., male holotype. 10 – genitalia; 11 – F9; 12 – F2; 13 – pedicel and F1; 14 – mouth parts and palpi; 15 – tarsal claw; 16 – wing. Scale bars =