Verbasceae

Mourad, Magdy M., 2015, Diversity and evolutionary trends in the floral characters of some taxa of Scrophulariaceae sensu lato, Adansonia 37 (1), pp. 149-159 : 157

In tribe Verbasceae

Verbascum letourneuxii , V. sinaiticum and V. sinuatum occupy primitive status due to the less advanced characters viz. five distinct sepals, sepal median, staminal and dorsal carpellary bundles from the central stele, five petals, dithecous anthers, five staminal traces originated from the central stele to five fertile stamens and numerous lateral carpellary bundles. Absence of nectariferous disc raises Verbascum letourneuxii over V. sinaiticum and V. sinuatum .

AT SUBFAMILY LEVEL

Antirrhineae

Antirrhineae acquired the most advanced position in Antirrhinoideae , this is in accord with Pennell (1935) and Thieret (1976), and disagrees with Schäferhoff et al. (2010). Tribe Gratioleae acquired an advanced position in the present study, this agrees with Bentham & Hooker (1876) and Wettstein (1891), and disagrees with Pennell (1935) and Thieret (1976).