Tuxedo cruralis (Van Duzee)

SCHUH, RANDALL T., 2004, Revision of Tuxedo Schuh (Hemiptera: Miridae: Phylinae), American Museum Novitates 3435 (1), pp. 1-26 : 11-13

Tuxedo cruralis (Van Duzee)

Figures 1 , 2 , 4

Plagiognathus diversus cruralis Van Duzee, 1917: 283 (n.var.).

Psallus breviceps: Pinto, 1982: 102 (host record, phenology).

Tuxedo cruralis: Schuh, 2001: 252 (n.comb.).

HOLOTYPE: Plagiognathus diversus cruralis Van Duzee : Male: [ USA]: ‘‘ Tunnel Road , Alameda Co., Cal., # 1, 13­V­17, W. M. Giffard Collector’ ’. Deposited in the California Academy of Sciences.

DIAGNOSIS: Distinctive among Tuxedo species in having all of the femora entirely infuscate. Most similar to T. flavicollis and T. susansolomonae in the dark coloration of the antennae in the males, but those species with the femora pale at least basally; differing from susansolomonae by the partial to complete, pale, transverse fascia on the hemelytra posterior to the apex of the scutellum in that species; differing from flavicollis by that species having the base of the corium entirely dark.

DESCRIPTION: Male: Relatively small to medium­sized; total length 2.77–3.15, length apex clypeus–cuneal fracture 1.91–2.17, width pronotum 0.84–0.95. COLORATION (fig. 1): Background coloration brownish black, rarely lighter; basal one­third to onehalf of cuneus creamy white, corium frequently pale at extreme base, pale area sometimes more extensive, rarely entending onto basal area of clavus; all antennal segments black; labial segments 2 and 3 pale; all coxae pale, all femora and tibiae infuscate. SUR­ FACE AND VESTITURE (figs. 1, 4F): Pronotum weakly to moderately shining, remainder of dorsum weakly shining. STRUC­ TURE: Body weakly cylindrical in cross section, appearing somewhat flattened when viewed from above; head as in figure 4A, C; meso­ and metathoracic pleura as figure 4D; femoral trichobothria as in figure 4H; claws as figure 4E. GENITALIA (figs. 2, 4G): Body of vesica relatively slender; vesica with a single, bifid apical spine extending well beyond secondary gonopore and envelope of vesica, of form similar to that of T. bicinctus and T. drakei , but spine longer and distinctly curving, unlike condition in those species.

Female: Relatively small to mediumsized; total length 2.83–3.13, length apex clypeus–cuneal fracture 2.03–2.26, width pronotum 0.95–1.06. COLORATION (fig. 1): Infrequently, as in male, dorsum largely dark with contrasting pale base of cuneus and base of corium weakly pale; more frequently dorsum partially to largely pale, but with calli, posterior half of corium, and distal half of cuneus dark brown and contrasting with remainder. SURFACE AND VESTITURE: As in male. STRUCTURE (fig. 4B): As in generic description.

HOST: Collected most frequently breeding on Ceanothus spp. (Rhamnaceae) , less frequently on Cercocarpus spp. (Rosaceae) . Also known from Quercus spp. (Fagaceae) , Shepherdia sp. (Elaeagnaceae) , Prunus sp. (Rosaceae) , Salix sp. (Salicaceae) , and Simmondsia sp. (Simmondsiaceae) .

DISTRIBUTION: Southern British Columbia, Canada , in the north, south to northern Baja California, Mexico , and east to western Colorado and New Mexico .

DISCUSSION: As noted by Schuh (2001), this species was originally described as a variety of Plagiognathus diversus Van Duzee (= Europiella artemisiae (Becker)) by Van Duzee (1917). In an earlier paper Van Duzee (1914) used the name Psallus breviceps Reuter (1909) , apparently in reference to T. cruralis ; Van Duzee (1914) noted that Dr. Reuter apparently had two pale females before him when describing the species from Texas. Van Duzee’s usage of the name breviceps seems to have been based solely on his reading of the literature, yet his indication that Reuter had examined two pale females contradicts information in the original description, where Reuter makes explicit reference only to the attributes of males and indicates that he had only two examples of the species, from ‘‘Texas’’. After correspondence with Herbert Zettel and I. M. Kerzhner, I have not been able to locate the specimens examined by Reuter, which were indicated as being deposited in the Naturhistorisches Museum in Vienna. This does not mean that they do not exist, but only that their identity has not yet been established. So, I am treating the name Psallus breviceps Reuter as a nomen dubium until such time as its identity can be verified.

Pinto (1982) also used the name Psallus breviceps , this time in explicit reference to T. cruralis . Pinto’s usage was based on identifications provided by T. J. Henry. I am uncertain on whose ientification Henry’s concept was based, for during the course of this study I have not located any specimens bearing identification labels reading ‘‘ Psallus breviceps’’ written by Van Duzee, Henry, or any other author.

Even in the absence of Reuter’s specimens, it seems quite certain that application of the name Psallus breviceps Reuter to what I am calling Tuxedo cruralis is in error. After extensive searching of existing collections from the western United States, and after having done extensive collecting in all of the western states, including western Texas, I have found no evidence to indicate that T. cruralis occurs in Texas. Thus, I am treating the use of Psallus breviceps by Van Duzee (1914) and Pinto (1982) as misidentifications.

This is the most widely distributed of the Tuxedo species and has been collected on the widest range of hosts, although is breeds most consistently on Ceanothus spp. and Cercocarpus spp.

SPECIMENS EXAMINED: CANADA: British Columbia: Oliver, Vaseaux Lake, June 26, 1959, L. A. Kelton, 1♀ (CNC). MEXICO: Baja California Norte: 1.5 km W of Parque Sierra San Pedro Martir, 2090 m, April 25, 1985, R. T. Schuh and B. M. Massie, Ceanothus cuneatus (Rhamnaceae) , 33, 4♀ (AMNH). 12 mi SE of El Rosario, March 25, 1979, J. D. Pinto, Simmondsia sp. (Simmondsiaceae) , 83, 16♀ (UCR). 13 mi N of Punta Prieta, March 26, 1979, J. D. Pinto, 1♀ (UCR). 2 mi W of Las Encinas, 5200 ft, March 18, 1972, J. Powell, Ceanothus sp. (Rhamnaceae) , 13, 1♀ (UCB). 2 mi W of Las Encinas, Sierra San Pedro Martyr, 3600 ft, March 18, 1972, J. Doyen, Ceanothus sp. (Rhamnaceae) , 33, 10♀ (UCB). 6 mi E of Ojos Negros, April 16, 1983, D. K. Faulkner, 13, 1♀ (SDNH). 7 mi SE of Maneadero, 100 ft, March 25, 1973, J. Doyen, 23, 2♀ (UCB). 7 mi W of Parque Sierra San Pedro Martir, 1720 m, April 25, 1985, R. T. Schuh and B. M. Massie, Ceanothus cuneatus (Rhamnaceae) , 103, 10♀ (AMNH). 7 mi W of Parque Sierra San Pedro Martir, 1720 m, April 25, 1985, R. T. Schuh and B. M. Massie, Ceanothus tomentosus (Rhamnaceae) , 73, 5♀ (AMNH). La Mission, Rio San Miguel, February 27, 1982, Brown and Faulkner, 13, 3♀ (SDNH). Las Encinas, Sierra San Pedro Martyr, May 25, 1980, Brown and Faulkner, 33, 4♀ (SDNH). Rancho El Progreso, March 25, 1979, E. M. Fisher, 1♀ (UCR). USA: Arizona: Graham Co.: Graham Mountains, Forest Road 808, T9S R25E, 5000 ft, April 25, 1982, M. D. Schwartz, Ceanothus greggii (Rhamnaceae) , 23, 4♀ (AMNH). Maricopa Co. : Reavis Ranch Trail, 3600 ft, April 19, 1982, D. A. and J. T. Polhemus, Quercus turbinella (Fagaceae) , 73, 5♀ (JTP). Yavapai Co.: 1 mi E of Yarnell, April 29, 1981, D. A. and J. T. Polhemus, Cercocarpus betuloides (Rosaceae) , 303, 8♀ (JTP). California: Alameda Co.: Berkeley, April 6, 1972, H. L. Jones, Cercocarpus traskiae (Rosaceae) , 33, 3♀ (CAFA). Oakland, hills back of city, April 3, 1931, E. C. Zimmerman, 13, 5♀ (UCB). Alpine Co.: 4 mi SE of Markleesville, 8150 ft, June 19, 1966, W. Gagne, 13 (UCB). Butte Co.: Oroville, April 30, 1927, H. H. Keifer, Salix hindsiana (Salicaceae) , 203, 18♀ (CAS). Oroville, April 30, 1927, H. H. Keifer, Salix hindsiana (Salicaceae) , 13, 1♀ (CNC). Yankee Hill, May 8, 1928, H. H. Keifer, Ceanothus cuneatus (Rhamnaceae) , 23 (CAS). Calaveras Co.: Mokelumne Hill, April 17, 1930, R. L. Usinger, 63, 8♀ (UCB). Contra Costa Co.: Moraga, May 13, 1977, R. F. Denno, 13 (UCD). Humboldt Co.: 2 mi S of Ettersburg, May 22, 1976, J. Hafernik, 13 (UCB). 3 mi E of Benbow, May 20, 1976, R. Dietz and J. Powell, 43, 7♀ (UCB). Beatrice, June 21, 1959, Kelton and Madge, 153, 10♀ (CNC). Beatrice, May 30, 1940, B. P. Bliven, 6♀ (CAS). Blocksburg, June 11, 1933 – June 10, 1951, B. P. Bliven, 13 (CAS). Bridgeville, June 20, 1959, Kelton and Madge, 73 (CNC). Bridgeville, June 20, 1959, Kelton and Madge, Ceanothus sp. (Rhamnaceae) , 6♀ (CNC). Dinsmores, June 25, 1939, B. P. Bliven, 33 (CAS). Garberville, May 5, 1940, B. P. Bliven, 23 (CAS). McCann, June 19, 1959, Kelton and Madge, 1♀ (CNC). Redway, May 15, 1938, B. P. Bliven, 13, 3♀ (CAS). Samoa, June 6, 1937, B. P. Bliven, 63, 7♀ (CAS). Shively, June 21, 1959, Kelton and Madge, 43, 7♀ (CNC). Kern Co.: 2.5 mi NNE of Caliente, April 28, 1978, J. D. Pinto, 13 (UCR). 4.5 mi E of Onyx, April 21, 1982, J. D. Pinto and R. K. Velten, Ceanothus sp. (Rhamnaceae) , 4♀ (UCR). 5 mi N of Kernville, May 15, 1968, S. W. Earnshaw, 23, 4♀ (UCB). 7 mi W of Wofford Heights on Rt 155, 1520 m, June 26, 1999, May 18, 2000, M. D. Schwartz, Ceanothus integerrimus (Rhamnaceae) , 11♀ (CNC). Havilah, April 28, 1978, J. D. Pinto, 33 (UCR). Lake Isabella, Main Dam Campground, April 24, 1980, M. D. Schwartz and L. Russell, Ceanothus integerrimus (Rhamnaceae) , 2♀ (OSU). Tehachapi Mts., Antelope Canyon, 4800–5200 ft, May 15, 1976, J. D. Pinto, Quercus turbinella (Fagaceae) , 23 (UCR). Walker Pass on Rt 178, 1600 m, May 18, 2000, M. D. Schwartz, Ceanothus sp. (Rhamnaceae) , 143, 4♀ (CNC). Walker Pass, 5250 ft, May 30, 1981, J. T. Polhemus, 13 (JTP). Lassen Co.: Susanville, June 17, 1959, Kelton and Madge, 1♀ (CNC). Los Angeles Co.: Claremont, C. F. Baker, 13 (USNM). Devils Canyon, June 16, 1948, E. I. Schlinger, 13 (UCD). Malibu Canyon, February 27, 1966, 33 (LACM). Mt. Wilson trail, upper half, June 5, 1915, H. Morrison, 23 (USNM). Pasadena, June 5, 1909, Grinnell, 73, 7♀ (CAS). Marin Co.: Mill Valley, April 25, 1926, M. C. Van Duzee, 13, 1♀ (CAS). Phoenix Lake, May 14, 1927, H. H. Keifer, Ceanothus sorediatus (Rhamnaceae) , 13, 4♀ (CAS). Ross, April 28, 1918, E. P. Van Duzee, 13 (CAS). Ross, April 28, 1918, J. C. Bradley, 13, 3♀ (CU). Mendocino Co.: 15 mi W of Laytonville, June 15, 1972, D. Veirs, 13 (UCB). 3 mi N of Branscomb, May 30, 1980, A. E. Hajek, 13 (UCB). Hopland Experiment Station, 2500 ft, May 23, 1971, C. B. Phillip, 13 (CAS). Modoc Co.: Fandango Pass Summit, Warner Mts., 1890 m, July 3, 1979, R. T. Schuh and B. M. Massie, Cercocarpus ledifolius (Rosaceae) , 5♀ (AMNH). Monterey Co.: 6.5 mi S of Big Sur, April 30, 1977, A. J. Mayor, Ceanothus sp. (Rhamnaceae) , 83, 8♀ (UCR). Chew’s Ridge, 5100 ft, June 23, 1967, R. G. Denno, 13 (UCD). Palo Colorado Road, 8 mi SSE of Hiway 1, Ventural Wildlands Trail, April 20, 1980, M. D. Schwartz and L. Russell, Ceanothus cuneatus (Rhamnaceae) , 123, 18♀ (OSU). Napa Co.: 2 mi NNE of Anguin, N side of Howell Mt., 1300 ft, May 1, 1986, H. B. Leech, 1♀ (CAS). 7 mi S of Pope Valley, April 24, 1971, J. Marsh, 13 (UCD). Soda Creek, May 3, 1932, H. H. Keifer, Cercocarpus betuloides (Rosaceae) , 3♀ (CAFA). Nevada Co.: 8 mi S of Grass Valley, May 16, 1930, E. P. Van Duzee, 1♀ (CAS). Orange Co.: Santiago Canyon, April 2, 1935, E. L. Paddock, Ceanothus megacarpus (Rhamnaceae) , 143, 5♀ (USNM). Santiago Canyon, April 2, 1936 – April 14, 1935, E. L. Paddock, Ceanothus megacarpus (Rhamnaceae) , 23 (CNC). Riverside Co.: Menifee Valley (hills on W end), February 28, 1985 – April 20, 1978, J. D. Pinto, Ceanothus crassifolius (Rhamnaceae) , 513, 50♀ (UCR). Milepost 29 on Rt 243, April 21, 1980, M. D. Schwartz and L. Russell, Cercocarpus sp. (Rosaceae) , 73, 3♀ (OSU). North Fork of San Jacinto River, San Jacinto Mts., Hwy 74, February 29, 1976, A. J. Mayor, Ceanothus cuneatus (Rhamnaceae) , 73, 3♀ (UCR). NW of Murietta, road to Tenaja Fire Station, May 13, 1978, J. D. Pinto, 13 (UCR). San Jacinto Mountains, jct of Poppet Flat Rd and Rt 243, 1270 m, May 20, 2000, M. D. Schwartz, Cercocarpus sp. (Rosaceae) , 1♀ (CNC). Sacramento Co.: Fair Oaks, April 5, 1931, H. H. Keifer, 13 (CAFA). Santa Barbara Co. : Santa Cruz Island, Prisoners Harbor, February 5, 1979, J. Powell, Ceanothus insularis (Rhamnaceae) , 23, 2♀ (UCB). San Benito Co.: Hwy 25 S of Paicines, March 17, 1986, C. B. Barr, Ceanothus sp. (Rhamnaceae) , 23 (LSU). Pinnacles National Monument, April 6, 1951, E. J. Taylor, 83, 6♀ (UCD). Pinnacles National Monument, April 6, 1951, E. J. Taylor, Ceanothus cuneatus (Rhamnaceae) , 1♀ (CAFA). San Bernardino Co.: Cajon Pass, jct Rts I­15 and 138, 1030 m, May 2, 1985, R. T. Schuh and B. M. Massie, Cercocarpus betuloides (Rosaceae) , 20♀ (AMNH). jct of Black Canyon and Cedar Canyon Roads, 3000 ft, May 18, 1982, M. D. Schwartz, Prunus fasciculata (Rosaceae) , 153, 9♀ (AMNH). San Bernardino Mts. above Mountain Home, 440 m, May 11, 1978, R. T. Schuh and J. D. Pinto, 13 (AMNH). San Bernardino, March 6, 1973, E. L. Paddock, Ceanothus sp. (Rhamnaceae) , 13 (CAFA). San Diego Co.: 2 mi SE of Santa Ysabel , May 5, 1983, J. D. Pinto, Ceanothus sp. (Rhamnaceae) , 13, 2♀ (UCR). Cibbets Flat Campground on Kimball Creek Road, 1280 m, April 29, 1985, R. T. Schuh, Ceanothus greggii (Rhamnaceae) , 103, 45♀ (AMNH). Decanso Junction on st. rt. 79, May 16, 1982, M. D. Schwartz, Ceanothus integerrimus (Rhamnaceae) , 243, 24♀ (AMNH). Laguna Mountain Rd (Old Hwy 80) just SE of jct with Rt 8, 1180 m, May 21, 2000, M. D. Schwartz Prunus fasciculata , 1♀ (CNC). McCain Valley, May 3, 1980, Brown and Faulkner, 2♀ (SDNH). Mt. Laguna, June 21, 1963, J. Powell, 13 (UCB). No specific locality, April 6, 1913 – May 12, 1913, E. P. Van Duzee, 73, 20♀ (CAS). Otay Mountain, May 18, 1978, D. K. Faulkner, 63, 6♀ (SDNH). Pine Valley, 1190 m, April 29, 1985, R. T. Schuh and B. M. Massie, Ceanothus integerrimus (Rhamnaceae) , 153, 15♀ (AMNH). San Luis Obispo Co.: 5 mi NS of Santa Margarita , May 16, 1980, J. D. PInto, 13, 2♀ (UCR). Atascadero, April 23, 1951, R. M. Bohart, 13 (UCD). Nacimiento Dam, April 14, 1967, P. A. Rude, 13, 1♀ (UCB). Santa Barbara Co. : 3 mi N of San Marcos Pass, Los Padres Natl. Forest, March 21, 1996, A. R. Gillogly, Ceanothus sp. (Rhamnaceae) , 3♀ (TAMU). Los Prietos, March 14, 1967, P. A. Opler, 83, 11♀ (UCB). Santa Cruz Island, Avalon Canyon, March 31, 1968, J. Powell, 33 (UCB). Santa Cruz Island, Central Valley, April 23, 1976, J. D. Pinto, Cercocarpus sp. (Rosaceae) , 223, 8♀ (UCR). Santa Cruz Island, Pebbly Beach Canyon, March 31, 1968, P. A. Opler, 23, 2♀ (UCB). Santa Cruz Island, UC Station Headquarters, February 3, 1979, J. Powell, 23 (UCB). Santa Cruz Island, Upper Central Valley, 1000 ft, March 15, 1969, P. A. Opler, Ceanothus arboreus (Rhamnaceae) , 33 Santa Clara Co. : 5 mi E of San Jose, Alum Rock Park, May 16, 1962, J. Doyen, 13 (UCB). Americh Road, April 21, 1996, M. Isaak, 13 (USNM). South end of Mines Road, 2350 ft, April 23, 1972, H. B. Leech, Ceanothus cuneatus (Rhamnaceae) , 33, 1♀ (CAS). Stanford University Campus, April 29, 1928, R. L. Usinger, 2♀ (CNC). Santa Cruz Co. : 2 mi E of Soquel at Bates Creek, May 2, 1971, J. Doyen, 23 (UCB). Felton, June 12, 1952, W. H. Lange, 13 (UCD). Shasta Co.: 12 mi W of Whiskeytown, May 14, 1997, W. F. Chamberlin, 13, 2♀ (TAMU). 7 mi S of Cottonwood, May 1, 1971, P. W. Oman, 1♀ (OSU). Castle Crag, June 25, 1939, Schuh and Gray, 1♀ (OSU). Hat Creek, May 11, 1944, W. F. Chamberlin, Ceanothus sp. (Rhamnaceae) , 13 (TAMU). Siskiyou Co.: 1 mi SE of Bartle, June 8, 1974, J. Doyen, 13 (UCB). Tehama Co. : 0.3 mi N of Red Bluff, 304 ft, April 14, 1981, M. D. Schwartz, Quercus sp. (Fagaceae) , 13, 7♀ (AMNH). Red Bluff, Dog Island State Park, April 29, 1984, D. S. Chandler, 33, 1♀ (UN­ HAMP). Tulare Co.: 10 mi S of Fairview, May 2, 1964, C. A. Toschi, 13 (UCB). 10.3 mi N of Lemon Cove on J21, March 25, 1981, M. D. Schwartz, Ceanothus integerrimus (Rhamnaceae) , 163, 5♀ (AMNH). 5.7 mi N of Kernville, Hospital Flat Campground, April 29, 1978, J. D. Pinto, Ceanothus sp. (Rhamnaceae) , 53, 5♀ (UCR). 9 mi S of Fairview, April 30, 1964, R. L. Langston, 23, 2♀ (AMNH). Potwisha, 3 mi NE Ash Mountain Headquarters, May 3, 1979, J. A. De­ Benedictis, 13 (UCB). Sequoia National Park, May 29, 1949, W. D. Pierce, Ceanothus sp. (Rhamnaceae) , 3♀ (LACM). Tuolumne Co.: 10 mi E of Buck Meadows, May 11, 1997, W. F. Chamberlin, 13, 1♀ (TAMU). Sonora, May 23, 1930, E. P. Van Duzee, 13 (CAS). Ventura Co.: 2 mi E of Lake Sherwood, March 16, 1967, P. A. Opler, 13 (UCB). Casitas Reserve, N end, March 15, 1967, P. A. Opler, 13, 5♀ (UCB). Ojai, April 24, 1935, C. E. Norland, 13 (LACM). Westlake Village, May 16, 1964, E. L. Paddock, 1♀ (CAFA). Yolo Co.: Putah Canyon, April 3, 1960, D. R. Miller, 13 (UCD). Yuba Co.: Dry Creek, Spenceville Wildlife Area, May 6, 1980, J. A. Powell, 23 (UCB). Colorado: Mesa Co. : Colorado National Monument, Liberty Cap Trailhead, 6000 ft, June 10, 1982, M. D. Schwartz, Cercocarpus montanus (Rosaceae) , 73, 51♀ (AMNH). Nevada: Elko Co.: Ruby Mts., Lomoille Canyon, E of Powerhouse Picnic Area, 6000 ft, June 16, 1983, R. T. Schuh and M. D. Schwartz, Cercocarpus ledifolius (Rosaceae) , 83, 18♀ (AMNH). Washoe Co.: Verdi, July 9, 1967, W. Gagne, 13 (UCD). White Pine Co.: 8.3 mi N of Hwy 50 on Steptoe Creek Rd, 7580 ft, July 19, 1980, G. M. Stonedahl, Cercocarpus ledifolius (Rosaceae) , 20♀ (OSU). Wheeler Peak Road W of Baker, Humboldt Natl. Forest, 2609 m, July 14, 1980, R. T. Schuh and G. M. Stonedahl, Cercocarpus ledifolius (Rhamnaceae) , 443, 81♀ (AMNH). New Mexico: Sandoval Co.: East Fork Jemez River, La Concha Campground, 8300 ft, August 14, 1972, R. E. Orth, 13 (UCR). Oregon: Harney Co.: 11 mi E of Frenchglen, 6000 ft, July 16, 1957, J. D. Lattin, Cercocarpus sp. (Rosaceae) , 1♀ (OSU). Josephine Co.: 12 mi W of Cave Junction, June 12, 1979, R. T. Schuh, Ceanothus integerrimus (Rhamnaceae) , 1♀ (AMNH). Wheeler Co.: 4.5 mi S of Mitchell on Summit Prairie Road, June 22, 1979, M. D. Schwartz, Cercocarpus ledifolius (Rosaceae) , 53, 15♀ (OSU). 4.5 mi S of Mitchell on Summit Prairie Road, June 22, 1979, R. T. Schuh, Cercocarpus ledifolius (Rosaceae) , 133, 13♀ (AMNH). Utah: Cache Co.: 5 mi NE of Logan, Green Canyon (trail), T12N R2E Sec. 21, 6000–6500 ft, July 3, 1982, M. D. Schwartz, Cercocarpus ledifolius (Rosaceae) , 63, 7♀ (AMNH). Green Canyon Trail, 6500 ft, June 27, 1980, M. D. Schwartz, Cercocarpus betuloides (Rosaceae) , 53, 8♀ (OSU). Carbon Co.: 8 mi NE of Helper, Price Canyon Rec. Area, T12E R9E, 8000 ft, July 9, 1982, M. D. Schwartz, Cercocarpus ledifolius (Rosaceae) , 33, 1♀ (AMNH). San Juan Co.: 27 mi E of Blanding at MP 101 on Rt 95 near Mule Tower Ruins, 6500 ft, June 12, 1982, M. D. Schwartz, Shepherdia rotundifolia (Elaeagnaceae) , 23 (AMNH). Natural Bridges National Monument (at campground), 6500 ft, June 17, 1983, R. T. Schuh and M. D. Schwartz, Shepherdia rotundifola (Elaeagnaceae) , 15♀ (AMNH). Natural Bridges National Monument, May 29, 1985, J. T. and D. A. Polhemus, Shepherdia canadensis (Elaeagnaceae) , 163, 17♀ (JTP). Natural Bridges National Park, T 36, 37E R 17, 18E, 6300 ft, June 12, 1982, M. D. Schwartz, Cercocarpus montanus (Rosaceae) , 11♀ (AMNH). Sevier Co.: 2.3 mi N of I­70 on road to Kanosh, 6980 ft, July 16, 1980, G. M. Stonedahl, Cercocarpus ledifolius (Rosaceae) , 2♀ (OSU). Dog Spring Rd at Rt 25, 2719 m, July 16, 1980, R. T. Schuh and G. M. Stonedahl, Cercocarpus ledifolius (Rhamnaceae) , 113, 23♀ (AMNH). Jct Mytoge Mtn Rd and Hwy 25, 8850 ft, July 16, 1980, G. M. Stonedahl, Cercocarpus ledifolius (Rosaceae) , 6♀ (OSU). Uintah Co.: 5–10 mi SW Bonanza, 5000– 5600 ft, May 14, 1981, M. D. Schwartz, 1♀ (AMNH).

Figures

Fig. 1. Habitus views of Tuxedo spp. See appendix 1 for localities.

Fig. 2. Male genitalia of Tuxedo spp. See appendix 1 for localities.

Fig. 4. Scanning electron micrographs of T. cruralis from USA: White Pine Co.: Wheeler Peak Road. A. Head, 3, lateral view. B. Head, ♀, lateral view. C. Head, 3, anterior view. D. Mesothoracic spiracle and metaepisternal scent efferent system, lateral view. E. Pretarsus, frontal view. F. Setae on hemelytra. G. Genital segment, posterior view of left paramere, phallotheca, and right paramere. H. Trichobothria on hind femur, lateroventral view. Scales as indicated.