Tipula (Vestiplex) virgatula virgatula Riedel, 1913

Starkevich, Pavel, Podenas, Sigitas & Gelhaus, Jon K., 2020, Taxonomic review of Tipula (Vestiplex Bezzi) crane flies (Diptera: Tipulidae) in Mongolia, Zootaxa 4837 (1), pp. 1-88 : 77-84

Tipula (Vestiplex) virgatula virgatula Riedel

( Figs 12 , 14 , 174–184 ; Map 14 )

Tipula virgatula Riedel, 1913: 58 .

Tipula subcarinata Alexander, 1922: 347 .

Tipula (Vestiplex) subcarinata: Alexander, 1935: 119 ; 1936a: 2.

Tipula (Vestiplex) subcarinata subcarinata: Savchenko, 1960: 192 ; Savchenko, 1964: 211 (synonymy).

Tipula (Vestiplex) virgatula virgatula: Savchenko, 1964: 211 .

Tipula edentata Alexander, 1922: 349 .

Tipula (Vestiplex) edentata: Savchenko, 1960 ; Savchenko, 1964: 211 (possible synonymy of T. (V.) virgatula ); Oosterbroek & Theowald, 1992: 160 (synonymy not certain); Podenas et al., 2013: 113 (as synonym of T. (V.) virgatula ).

Type material examined. Tipula (Vestiplex) virgatula Riedel : HOLOTYPE ♂: MONGOLIA, N. Mongolei Leder 92, Tipula virgatula , m. [mihi] ♂, Type det. M.P. Riedel, Typ ( NHMV). COTYPE: ♂, N. Mongolei Leder 92, Tipula virgatula m. [mihi] ♂, Type det. M.P. Riedel, Cotype Riedel = T. (Vestiplex) subcarinata Alexander [identified by E.N. Savchenko] ( ZMHB).

Tipula (Vestiplex) edentata Alexander : HOLOTYPE ♀: MONGOLIA, August 1909, Dr. du Chazaud, Mission de Lacoste ( MNHN). Wing slide mounted ( USNM).

Note on types. According to Alexander (1922) the type specimen of T. (V.) edentata is preserved at MNHN, however its presence in the collection is not confirmed .

Additional material examined. MONGOLIA. Arkhangay Aimag: 2 ♂, Tsenkher Soum, Nuuriin Khooloi lake ~ 9 km SW Tavanbulag, N47.33933, E101.82093, 1693 m, 10.vii.2004, SRPT, SRP 04071001; 3 ♀, Tsenkher Soum, Tsetserleg Gol ~ 50 km SSW Tavanbulag, N47.04409, E101.76352, 1902 m, 11.vii.2004, SRPT, SRP 04071102; 24 ♂, 8 ♀, Ikhtamir Soum, NW braid of Khoit Tamir Gol ~ 25 km SW Ikhtamir, N47.50523, E100.93208, 1728 m, 14–15.vii.2004, SRPT, SRP 04071402; 15 ♂, 2 ♀, Ikhtamir Soum, Khoit Tamir Gol ~ 29 km SW Ikhtamir, N47.48567, E100.87875, 1749 m, 15.vii.2004, SRPT, SRP 04071501; 11 ♂, 2 ♀, Chuluut Soum, Khanui Gol ~ 60 km SW Ikhtamir, N47.50949, E100.57510, 1981 m, 15.vii.2004, SRPT, SRP 04071502; 12 ♂, 1 ♀, Chuluut Soum, Khurmen/Davaat Gol 15 km SSE Chuluut/Jargalant, N47.42580, E100.30130, 2104 m, 15–16.vii.2004, SRPT, SRP 04071503; 2 ♂, Chuluut Soum ponds at Egiin Davaa ~ 47 km SW Chuluut/Jargalant, N47.21198, E99.91114, 2582 m, 16.vii.2004, A.E.Z. Short, J.K. Gelhaus, Enkhnasan, SRP 04071601; 1 ♂, Chuluut Soum, Tsohiotiin Davaa (mountain pass), N47.68999, E100.21012, 2256 m, 17.vii.2004, J.K. Gelhaus #969; 1 ♂, Ikhtamir Soum, E side of Khoit Tamir Gol 4 km NE Ikhtamir, N47.59928, E101.24521, 1602 m, 14.vii.2004, SRPT, SRP 04071401; 19 ♂, Chuluut Soum, Chuluutin Gol 40 km N Chuluut/Jargalant, N47.890440, E100.31810, 1937 m, 17.vii.2004, SRPT, SRP 04071701; 88 ♂, 4 ♀, Tariat Soum, Nariin Gichgenii Gol/Urd Terkh Gol ~ 5 km SE Khorgo/Tariat, N48.12661, E99.94128, 2019 m, 17–18.vii.2004, SRPT, SRP 04071702; 67 ♂, 3 ♀, Tariat Soum, Terkhiin Tsagan Nuur ~ 8 km W of Khorgo/Tariat, N48.14404, E99.78300, 2057 m, 18.vii.2004, SRPT, SRP 04071801; 88 ♂, 6 ♀, Tariat Soum, Khunt Nuur ~ 3 km SW Tsagaan nuur Brigade, N48.07248, E99.52216, 2062 m, 18.vii.2004, SRPT, SRP 04071802. Bayan Olgii Aimag: 3 ♂, Tsengel Soum, Sul Uuliin Gol, trib. of Tsagaan Gol, Altai-Tavan Bogd NP, N49.10448, E88.04845, 2796 m, 10.vii.2008, S. Podenas, MAIS 08071001; 28 ♂, 3 ♀, Tsengel Soum, Tsagaan Gol, 20 km W Zagast Nuur Bag, Entrance to Altai-Tavan Bogd NP, N49.09161, E88.10436, 2410 m, 12.vii.2008, S. Podenas, MAIS 08070902; 3 ♂, Tsengel Soum, Mogoitiin Gol 13 km ESE Asgat Uul, N48.75859, E88.60339, 2151 m, 14.vii.2008, S. Podenas, MAIS 08071305; 56 ♂, 11 ♀, Tsengel Soum, Khoton Nuur 15 km NW Syrgal, N48.66722, E88.29927, 2086 m, 14–15.vii.2008, S. Podenas, R.W. Bouchard, Jr., MAIS 08071402; 1 ♂, Tsengel Soum, Partizanii Bulag below Khazabai Sala (waterfall), 2148 m, N48.61734, E88.31315, 16.vii.2008, R.W. Bouchard, Jr., MAIS 08071602; 1 ♂, 1 ♀, Tsengel Soum, Syrgali Gol, bridge over outflow between Khoton Nuur and Khurgen Nuur at Syrgal, 2095 m, N49.59925, E88.43707, 16.vii.2008, Suvdaa, MAIS 08071603; 1 ♂, Tsengel Soum, Sumdairag Gol, 12 km S Syrgai, 2129 m, N48.50679, E88.50977, 16.vii.2008, J. Morse, Suvdaa, S. Podenas, MAIS 08071604; 20 ♂, 6 ♀, Tsengel Soum, Rashaanii Gol, at bridge, trib. of Tsagaan Us Gol, N48.75565, E88.15266, 2098 m, 15.vii.2008, S. Podenas, MAIS 08071501; 15 ♂, Tsengel Soum, Khar Ovoo Gol, draining Khelkhee Lake, N48.62918, E88.28554, 2147 m, 15–16.vii.2008, MAIS 2008071601; 2 ♂, Sagsay Soum, Yamaatiin Gol, 17 km SW Dayan Military Base, N48.16728, E88.85435, 2073 m, 17.vii.2008, MAIS 2008071703; 1 ♂, Sagsay Soum, pools attached to Dayan Lake, 2km N Dayan Military Base, N48.27453, E88.86900, 2230m, 18.vii.2008, MAIS 2008071801; 3 ♂, Deluun Soum, Burgedtein Gol 3.5 km S Buyant Gol, N47.58121, E91.17047, 2049 m, 4.vii.2009, Mountainous stream with willows on banks, grass short, grazed; 2 ♂, Nogoonnuur Soum, Zakhin Us Gol, 15 km W Nogoonnuur, N49.57774, E90.03497, 1764 m, 8–9.vii.2010, MAIST, MAIS 2010070803; 1 ♂, Nogoonnuur Soum, unnamed ponds, 20 km WNW Nogoonnuur, Siylkhemiyn Nuruu Protected Area, N49.71056, E89.98618, 1751 m, 10.vii.2010, MAIST, MAIS 2010071001. Bayankhongor Aimag: 5 ♀, Changaj Mountains, Ulaan colon, 18 km S Pass Egijn davaa, 2300 m, Exp. Dr. Z. Kaszab, 1966, Nr. 552, 21.vi.1966 ( HNHM); 13 ♂, Erdenesogt Soum, Shargaljuut Gol, 10 km NE Shargaljuut Rashaan Hot Spring, jct Ugalzyn Gol, N46.35902, E101.27559, 2186 m, 4.vii.2011, MAIS 2011070402; 1 ♂, Erdenesogt Soum, Erkhetkhayrkhan Gol, crossing to Khokh Gol, N46.62626, E101.36736, 2500 m, 5.vii.2011, S. Podenas, MAIS 2011070501. Bulgan Aimag: 12 ♂, 3 ♀, Bulgan Soum, Urd Tamir Gol ~ 38 km SW Tsetserleg, N47.28244, E101.18793, 1872 m, 12–13.vii.2004, SRPT, SRP 04071201; 5 ♂, Bulgan Soum, unnamed hillside trib. of Khairkhan Davaani Gol, N46.94286, E100.85532, 2311 m, 13.vii.2004, J.K. Gelhaus, J.C. Morse, A.E.Z. Short, SRP 04071303; 1 ♂, Bulgan Soum, Urd Tamir Gol braid upstream of bridge ~ 63 km SW Tsetserleg, N47.11192, E101.01048, 2066 m, 13–15.vii.2004, SRPT, SRP 04071302; 1 ♂, Teshig Soum, wetland near confluence of Tariakhtain Gol and Egiin Gol, N49.74339, E103.10503, 936 m, 10.vii.2005, SRPT, SRP 05071002; 41 ♂, Teshig Soum, Egiin Gol, 12.7 km SW Teshig, N49.88872, E102.50978, 998 m, 11–13.vii.2005, SRPT, SRP 05071101; 12 ♂, 1 ♀, Teshig Soum, wetland, 4.3 km SSW Teshig, N49.90725, E102.65665, 970 m, 10– 11.vii.2005, SRPT, SRP 05071003; 2 ♂, Bureghangay Soum, steppe, grassland along roadside, 63 km SE Orhon, 1028 m, N48.26890, E104.19501, 26.vi.2006, J.K. Gelhaus #1047; 1 ♂, Birch ( Betula )-larch ( Larix ) forest, 15 km NW Bulgan, 1435 m, N48.89933, E103.36507, 26.vi.2006, J.K. Gelhaus #1048. Darkhan Aimag: 1 ♂, Shariingol Soum, Khuitnii Gol, N49.46450, E106.66871, 852 m, 21.vii.2003, SRPT, SRP 03072103. Dornod Aimag: 3 ♂, Bayn Uul Soum, pine forest, 2 km S Ulikhan Deed Ulikhan town, N49.49401, E112.56936, 853 m, 21.vii.2011, S. Podenas, MAIS 2011072101. Hovsgol Aimag: 6 ♂, Hovsgol lake, Borsog Gol (river) at shoreline, 1650 m, 3.vii.1996, E. Hunter #96031; 7 ♂, Khankh Soum, Sevsuul River, N51.1623611, E100.753833, 1656 m, 25.vi.2005, SRPT; 1 ♂, Khankh Soum, Noyon River, N51.2029722, E100.805833, 1707 m, 25.vi.2005, SRPT; 1 ♀, Khankh Soum, Shagnuul River, N51°15′18.6″, E100°50′45.8″, 1728 m, 25.vi.2005, SRPT; 9 ♂, Khankh Soum, Turag River, N51°17′11.1″, E100°49′12.3″, 1665 m, 25–26.vi.2005, SRPT; 3 ♂, Khankh Soum, mouth of Turag River—Hovsgol lake, N51°18′17.2”, E100°47′23.2″, 1655 m, 26.vi.2005, SRPT; 1 ♂, Khankh Soum, Turag River, N51°17′12.6″, E100°49′49.0″, 1663 m, 26.vi.2005, SRPT; 13 ♂, Khankh Soum, Turag River, N51°17′28.1″, E100°48′49.8″, 1668 m, 26.vi.2005, SRPT; 1 ♂, Khankh Soum, Turag River, N51°19′33.7″, E100°54′05.7″, 1795 m, 26.vi.2005, SRPT; 17 ♂, 1 ♀, Khankh Soum, Shagnuul River and Hovsgol lake nearby, N51°15′46.8″, E100°46′44.7″, 1657 m, 27.vi.2005, SRPT; 18 ♂, Khankh Soum, Shagnuul Gol, 1694 m, 27.vi.2005, SRPT; 1 ♂, Khankh Soum, Shagnuul Gol, 1727 m, 27.vi.2005, SRPT; 1 ♂, Khankh Soum, mouth of Noyon Gol at Hovsgol lake, N51°12′47.3″, E100°45′11.1″, 1607 m, 28.vi.2005, SRPT; 3 ♂, Khankh Soum, Sevsuul Gol and shore of Hovsgol lake, N51°09′42.0″, E100°43′42.2″, 1658 m, 28.vi.2005, SRPT; 1 ♂, Khankh Soum, Sevsuul Gol, N51°09′37.1″, E100°45′31.3″, 1666 m, 29.vi.2005, SRPT; 1 ♂, Khankh Soum, Sevsuul Gol, N51°09′22.3″, E100°46′36.6″, 1686 m, 29.vi.2005, SRPT; 3 ♂, Khankh Soum, Dalbay River, N51.00271, E100.49329, 1726 m, 1.vii.2005, SRPT; 3 ♂, Erdene Bulgan Soum, Emt Gol, 33.2 km NW Tarialyn, N49.87742, E101.82073, 1496 m, 13–14.vii.2005, SRPT, SRP 05071302; 1 ♂, Chandmani-Ondor Soum, Hohoo Gol, 1.5 km SE Chandmani-Ondor, N50.46416, E100.9491, 1251 m, 18.vii.2005, SRPT, SRP 05071801; 1 ♂, Chandmani-Ondor Soum, west branch (unnamed) of Khalkhan Gol, upstream of Bolnain Rashaan, 58 km NE Hatgal, N50.78323, E100.79572, 1676 m, 18–19.vii.2005, SRPT, SRP 05071803; 2 ♂, east trib. of upper Hohoo Gol (Bugat Gol), 44.5 km NE Hatgal, N50.69174, E100.66691, 1595 m, 19.vii.2005, SRPT, SRP 05071902; 9 ♂, Bayanzurkh Soum, Altargana Gol, 4.2 km N Bayanzurkh, N50.21297, E98.96367, 1662 m, 21–22.vii.2005, SRPT, SRP 05072103; 9 ♂, Khankh Soum, Dalbay valley, N51.0240556, E100.7558611, 1670 m, 24–25.vi.2005, SRPT; 2 ♂, Tumurbulag Soum, Bugcei (Bugceiin) Gol, 45.5 km SW Moron, N49.26168, E99.88242, 1618 m, 23.vii.2005, SRPT, SRP 05072301; 1 ♂, Tsagaan-Uur Soum, Uilgan Gol, 26 km ENE Tsagaan-Uur, N50.6231, E101.87375, 1152 m, 15–16.vii.2005, SRPT, SRP 05071502; 1 ♂, Tumurbulag Soum, N49.23175, E99.90027, 1790 m, 23.vii.2005, SRPT, SRP 05072302; 11 ♂, Hovsgol Aimag Steppe grassland; 23 km ESE Moron, 1554 m, N49.57650, E100.46652, 27.vi.2006, J.K. Gelhaus #1050; 2 ♂, Arbulag Soum, Ovor Gylaadin Gol (pooled stream), 7.0 km SE Sumber, 1987 m, N50.12844, E99.60749, 27–28.vi.2006, J.K. Gelhaus #1051, SRP 06062701; 9 ♂, Bayanzurh Soum, Ovor Gylaadin Gol (small stream), 20.0 km NW Sumber, 1802 m, N50.25681, E99.28587, 28.vi.2006, J.K. Gelhaus #1052, SRP 06062801; 5 ♂, 7 ♀, Hovsgol Aimag Ulaan-Uul Soum, Beltes Gol (river) 30.0 km NE Bayanzurh, 1791 m, N50.31142, E99.32278, 28.vi.2006, J.K. Gelhaus #1053, SRP 06062802; 2 ♂, Ulaan-Uul Soum, unnamed trib. (stream) of Beltes Gol, 34.0 km NE Bayanzurh, 2113 m, N50.42980, E99.21690, 28.vi.2006, J.K. Gelhaus #1054-A, SRP 06062803; 1 ♂, Hovsgol Aimag Ulaan-Uul Soum, Guna (Gunain) Gol (river), 51 km NE Bayanzurh, 1761 m, N50.61893, E99.12289, 29.vi.2006, J.K. Gelhaus #1055, SRP 06062903; 4 ♂, Ulaan Uul Soum, larch forest on hilltop, 15 km NNE Ulaan Uul, 1704 m, N50.81465, E99.32621, 29.vi.2006, J.K. Gelhaus #1056; 5 ♂, 1 ♀, Ulaan Uul Soum, Bagtag Gol (river) and wetland ponds/marshy meadow, 15 km NNE Ulaan Uul, 1598 m, N50.80484, E99.32228, 29–30.vi.2006, J.K. Gelhaus #1057, SRP 06062904 & 06063001; 1 ♂, 1 ♀, Renchinlhumbe Soum, Hog Gol (river), 25 km W Renchinlhumbe, 1569 m, N51.09576, E99.31911, 1.vii.2006, J.K. Gelhaus #1060, SRP 06070101; 8 ♂, Renchinlhumbe Soum, wetland (3 ponds) in sandy grassland, 19 km N Renchinlhumbe, 1595 m, N51.27236, E99.71152, 2.vii.2006, J.K. Gelhaus #1063, SRP 06070202; 2 ♂, Renchinlhumbe Soum, Jaray (Jarin) Gol (river), 33 km N Renchinlhumbe, 1580 m, N51.39865, E99.75060, 2– 3.vii.2006, J.K. Gelhaus #1064, SRP 06070203; 31 ♂, 2 ♀, Renchinlhumbe Soum, Outlet stream of Targan Nuur (lake), ~ 11 km NE Tsagaannuur (town), 1552 m, N51.42940, E99.42113, 3.vii.2006, J.K. Gelhaus #1065, SRP 06070301; 2 ♂, Tsagaannuur Soum, unnamed trib. of Shishged Gol ~ 16 km NW Tsagaannuur (town), 1606 m, N51.45833, E99.20051, 3–4.vii.2006, J.K. Gelhaus #1066, SRP 06070302; 4 ♂, Renchinlhumbe Soum, Larch ( Larix siberica ) forest, 41 km S Tsagaannuur (town), 1604 m, N50.98796, E99.23112, 4.vii.2006, J.K. Gelhaus #1071; 2 ♂, 3 ♀, Ulaan-Uul Soum, Guna (Gunain) Gol (river), 11 km NNW Ulaan-Uul, N50.77518, E99.18892, 1615 m, 4.vii.2006, SRP 06070403; 1 ♂, Bayanzurh Soum, Beltes Gol (river), 10 km E Bayanzurh, 1688 m, N50.20129, E99.08625, 5.vii.2006, J.K. Gelhaus #1072; 22 ♂, Bayanzurh Soum, Beltes Gol (river), 6 km ENE Bayanzurh, 1682 m, N50.19880, E99.03148, 6.vii.2006, SRP 06070601; 10 ♂, Bayanzurh Soum, Emtin Gol (stream), ~ 2 km S Bayanzurh, 1625 m, N50.15020, E99.97738, 6.vii.2006, J.K. Gelhaus #1075, SRP 06070602; 11 ♂, 1 ♀, Moron Soum, Tunamal Nuur (lake), 5.5 km W Arbulag, 1871 m, N49.89920, E99.39433, 6–7.vii.2006, J.K. Gelhaus #1076, SRP 06070603; 2 ♂, Ikh Uul Soum, Selenge Moron (river) ~ 8 km W Ikh-Uul, 1069 m, N49.44641, E101.33625, 7–8.vii.2006, J.K. Gelhaus #1077, SRP 06070701. Khentii Aimag: 1 ♂, Khovd Aimag, Khovd Soum, Dogd Gol, 3 km W Khovd soum centre, 1745 m, N48.12139, E91.33124, 1.vii.2008, S. Podenas, MAIS 08070102; 3 ♂, Khovd Aimag, Duut Soum. Tsagaan Burgasii Gol ~ 15 km NE Duut, N47.55936, E91.76095, 1865 m, 18–19.vii.2009; 1 ♂, Khovd Aimag, Duut Soum, Nevtiin Rashaan Springs jct Tsagaanburgasii Gol 9 km E Duut, N47.53136, E91.70514, 2051 m, 19.vii.2009; 4 ♂, Omnodelger Soum, Khurkhiyn Gol, 39 km NE Tsenkhermandal, N48.04989, E109.32549, 1364 m, 15.vii.2011, S. Podenas, MAIS 2011071502; 1 ♂, Binder Soum, Onon Gol, jct Khurkh Gol, 6.5 km SE Binder, N48.58405, E110.67959, 1039 m, 17.vii.2011, S. Podenas, MAIS 2011071701; 2 ♂, Binder Soum, Khurkh Gol, jct Onon Gol, 6.5 km SE Binder, N48.58354, E110.67884, 1038 m, 17.vii.2011, MAIS 2011071702; 4 ♂, Dadal Soum, Uuts Gol, 30 km NE Dadal, N49.24091, E111.84976, 904 m, 19.vii.2011, S. Podenas, MAIS 2011071902; 1 ♂, Norovlin Soum, Ulz Gol, 45 km SW Bayan Uul, N48.86579, E112.20216, 963 m, 21.vii.2011, S. Podenas, MAIS 2011072102. Ovorhangay Aimag: 3 ♂, Changaj Mountains, 21 km E somon Narijnteel, 2080 m, Exp. Dr. Z. Kaszab, 1964, Nr. 213, 27.vi.1964 ( HNHM); 2 ♂, Changaj Mountains, 5 km NE somon Chusirt, 1730 m, Exp. Dr. Z. Kaszab, 1964, Nrs. 225, 226, 29.vi.1964 ( HNHM); 20 ♂, Esonzyil Soum, Zegst/Sarin Gol, 74 km NW Arvayheer, N46.79062, E103.32486, 1665 m, 4–5.vii.2004, SRPT, SRP 04070403; 6 ♂, Kharkhorin Soum, Sant Uul pass, 23 km SE Shinebrigade, N46.87219, E103.14081, 2007, 5.vii.2004, SRPT, SRP 04070501; 1 ♂, Zuunbayan-Ulaan Soum, Millenium Road A0301 marker 380.6 km ~ 50 km NE Arvayheer, N46.60919, E103.10400, 1820 m, 5. vii.2004, SRPT, SRP 04070502; 43 ♂, 7 ♀, Hujirt Soum, Khavtsaliin Gol valley, N46.76292, E103.02561, 1750 m, 5–6.vii.2004, SRPT, SRP 04070504; 4 ♂, Khujirt Soum, Kharzanii Gol 12 km SE Khujirt, N46.80020, E102.84406, 1706 m, 6.vii.2004, SRPT, SRP 04070601; 7 ♂, Ovporhangay Aimag, Bat-Olziy Soum\ Orkhon Gol ~ 34 km W Khujirt, N46.89303, E102.39457, 1610 m, 6.vii.2004, SRPT, SRP 04070602; 35 ♂, 2 ♀, Bat-Olziy Soum, braid of Orkhon Gol ~ 40 km W Khujirt, N46.88586, E102.34386, 1646 m, 6–7.vii.2004, SRPT, SRP 04070603; 7 ♂, Bat-Olziy Soum, Orkhon’s Waterfall on Ulaan Gol ~ 300 m S Orkhon Gol, ~ 84 km W Khujirt, N46.78742, E101.96021, 1809 m, 7.vii.2004, SRPT, SRP 04070701; 107 ♂, 39 ♀, Bat-Olziy Soum, Ulaan Gol ~ 93 km W Khujirt, N46.73093, E101.88583, 1894 m, 7–8.vii.2004, SRPT, SRP 04070702; 1 ♂, 1 ♀, Kharkhorin Soum, Kharzani Gol 3.5 km NE Shankh Bridgade, N47.05722, E102.99191, 1519 m, 8.vii.2004, SRPT, SRP 04070803; 3 ♂, Kharkhorin Soum, Orkhon Gol 6 km W Kharkhorin center, N47.20226, E102.80169, 1462 m, 8–9.vii.2004, SRPT, SRP 04070901; 5 ♂, Bat-Olziit Soum, Orkhon gol, N46.79082, E101.93123, 1819 m, 24.vi.2011, O. Yadamsuren, MAIS 2011062401; 1 ♂, Ovorkhangay Aimag, Olziyt Soum, E Sanguiyn Dalay Nuur, N46.67575, E103.30890, 1707 m, 2.vii.2011, S. Podenas, MAIS 2011070202. Selenge Aimag: 4 ♂, Mandal Soum, Boyant Gol (trib. of Tunkhel Gol), 13.8 km above Tunkhel, 2.0 km upstream of Shonkhor Camp, N48.67237, E106.87851, 1078 m, 16–17.vii.2003, SRPT, SRP 03071602; 1 ♂, Javkhlant Soum, wetland/pools near left river bank, Yeroo River, 11.5 km upstream of Dulaan Khaan bridge, N49.84014, E106.33228, 647 m, 22–23.vii.2003, SRPT, SRP 03072202; 1 ♂, Khuder Soum, Khandgait Lake, 35.4 km E Khandgait Brigade (farm) and c. 18 km SW Khuder, N49.72086, E107.32710, 819 m, 23.vii.2003, SRPT, SRP 03072301; 1 ♂, Baruunharaa Soum, Haraa Gol, 2.3 km S Bayangol, N48.87738, E106.12375, 807 m, 4–5.vii.2005, SRPT, SRP 05070402; 2 ♂, Zuunburen Soum, Selenge Gol, 7.8 km NW Zuunburen, N50.10285, E105.78217, 647 m, 5–6.vii.2005, SRPT, SRP 05070501; 3 ♂, Khotol Soum, Orkhon Gol, 15.0 km WNW Orkhon, N49.1633, E105.18749, 756 m, 26–27.vii.2005, SRPT, SRP 05072602; 1 ♂, Eroo Soum, Bayan gol, N49.6298, E107.01804, 820 m, 1.vii.2011, O. Yadamsuren, MAIS 2011070102; 1 ♂, Khuder Soum, Bayan gol, N49.61433, E106.92656, 751 m, 1.vii.2011, O. Yadamsuren, MAIS 2011070101. Tov Aimag: 7 ♂, Erdene Soum, Gorkhi Terelj NP, W Bayangin Gol 8.2 km upstream of Tuul River Road, N48.08026, E107.63841, 1527 m, 6.vii.2003, SRPT, SRP 03070603; 2 ♂, Erdene Soum, Gorkhi Terelj NP, Khokh Chuluutiin Gol, N48.12675, E107.57996, 1680 m, 6.vii.2003, SRPT, SRP 03070604; 2 ♂, Tov Aimag Erdene Soum, Gorkhi Terelj NP, E Bayangin Gol, 12.9 km upstream from Tuul River Bridge, N48.14846, E107.75838, 1596 m, 7.vii.2003, SRPT, SRP 03070703; 4 ♂, Erdene Soum, Gorkhi Terelj NP, E Bayangin Gol, 4.2 km upstream of Tuul River Bridge, N48.07956, E107.77729, 1526 m, 7.vii.2003, SRPT, SRP 03070704; 12 ♂, Erdene Soum, Gorkhi Terelj NP, Tuul River, 10.1 km upstream of Tuul River Bridge, N48.09549, E107.84265, 1531 m, 8.vii.2003, SRPT, SRP 03070801; 2 ♂, Erdene Soum, Gorkhi Terelj NP, unnamed braid of Tuul River, N48.14846, E107.91267, 1564 m, 8–9.vii.2003, SRPT, SRP 03070805; 12 ♂, 1 ♀, Erdene Soum, Gorkhi Terelj NP, unnamed trib. of Tuul River on E side 8.5 km downstream of Galtain Gol, N48.09720, E107.84928, 1542 m, 11.vii.2003, SRPT, SRP 03071101; 9 ♂, Cherlen gol, near Cherlen Gol River, N48.11688, E108.37866, 1393 m, 5–6.viii.2003, SRPT; 2 ♂, env. Ulaan Baatar, Ihtenger, Tuul gol, N47.88349, E106.98829, 23.vi.2006, S. Podenas, SRP 06062302; 1 ♂, Gorkhi Terelj NP, S Terelj, N47°56′24″, E107°27′06″, 1626 m, 28–29.v.2009, S. Podenas, MAIS 2009062802; 5 ♂, Gorkhi-Terelj NP, Tuul River, near park entrance, N47.82965, E107.35424, 1410 m, 30.vi.2010, J.K. Gelhaus #1287, C. R. Nelson, A. Reschikov, T.A. McKnight; 1 ♂, Erdene Soum, Gorkhi Terelj NP, stream 0.5 km N Gorkhi Davaa (Pass), N47.94981 E107.45511, 1593 m, 11.vii.2011, S. Podenas, MAIS 2011071101; 1 ♀, Mongonmorit Soum, Zuun Baydlag Gol, downstream, 15 km SW Mongonmorit, N48.10818, E108.36736, 1475 m, 14.vii.2011, S. Podenas, MAIS 2011071402. Ulaanbaatar: 3 ♂, Ulaanbaatar, city center nr. Nat. Univ. Mongol. bldg. 2, N47.92494, E106.92178, 1305 m, 28.vi.2010, MAIST. Uvs Aimag: 1 ♂, 1 ♀, Tarialan Soum, Kharkhiraa Gol, 4 km W Tarialan, N49.77959, E91.86026, 1453 m, 2–3.vii.2010, MAIST, MAIS 2010070201; 2 ♂, Tarialan Soum, Khartarvagatay Gol, 30 km SW Tarialan, N49.55006, E91.70007, 2309 m, 3.vii.2010, MAIST, MAIS 2010070301; 1 ♂, 1 ♀, Tarialan Soum, Nutsgen Gol, 70 km SW Tarialan, N49.53507, E91.60309, 2552 m, 3.vii.2010, MAIST, MAIS 2010070302; 2 ♂, Tarialan Soum, Dund Gol, 73 km SW Tarialan, N49.51831, E91.58729, 2464 m, 3.vii.2010, MAIST, MAIS 2010070303; 1 ♂, Omnogovi Soum, unnamed lake (1.5-2 km 2), 24 km NNE Omnogovi, N49.32741, E91.82642, 2069 m, 4.vii.2010, MAIST, MAIS 2010070401; 2 ♂, Khovd Soum, Goojuur Gol, 25 km NE Khovd, N49.42978, E91.19503, 1975 m, 5–6.vii.2010, MAIST, MAIS 2010070505; 5 ♂, Khovd Soum, Shiver/Ireg Gol, 25 km NE Khovd, N49.43119, E91.19093, 1964 m, 5– 6.vii.2010, MAIST, MAIS 2010070504; 2 ♂, Khovd Soum, Goojuur Waterfall, 29 km NE Khovd, N49.44144, E91.25202, 2274 m, 6.vii.2010, MAIST, MAIS 2010070601; 2 ♂, Khovd Soum, springs by Goojuur Gol, 27 km NE Khovd, N49.43095, E91.21487, 2274 m, 6.vii.2010, MAIST, MAIS 2010070602; 6 ♂, Bokhmoron Soum, Altan Gadasni Gol, 7 km N Jigertei Bag Gulzat Protected Area, N49.99679, E90.27887, 1763 m, 10–11.vii.2010, MAIST, MAIS 2010071003, J.K. Gelhaus #1323; 1 ♂, Sagil Soum, Tsagaan Gol, 10 km W Uureg Nuur Bag, Gulzat Protected Area, N50.17117, E90.72800, 1629 m, 11–13.vii.2010, MAIST, MAIS 2010071103; 1 ♂, Turgen Soum, Khondlon Gol, 4.5 km W Turgen, N50.08403, E91.61472, 1295 m, 13–14.vii.2010, MAIST, MAIS 2010071303; 7 ♂, Turgen Soum, springs S side of Khondlon Gol, 6 km W Turgen, N50.07458, E91.60140, 1316 m, 14.vii.2010, MAIST, MAIS 2010071401; 2 ♂, Davst Soum, Tokhilog Gol, near Mongolian military border outpost, N50.69165, E92.59283, 1003 m, 14–15.vii.2010, MAIST, MAIS 2010071403; 36 ♂, 1 ♀, Sagil Soum, Borshoo Gol, 29 km NNE Sagil, N50.57615, E91.77029, 1281 m, 15–16.vii.2010, MAIST, MAIS 2010071502; 1 ♂, Turgen Soum, Javartain Gol, 33.5 km SW Turgen, N49.88653, E91.34708, 1871 m, 16.vii.2010, MAIST, MAIS 2010071603; 22 ♂, Turgen Soum, Javartain Gol & Turgen Gol, 33 km SW Turgen, N49.89234, E91.35239, 1849 m, 16–17.vii.2010, MAIST, MAIS 2010071602; 1 ♂, Turgen Soum, Buural Gol, 30 km SW Turgen, N49.90381, E91.41097, 1812 m, 17.vii.2010, MAIST, MAIS 2010071605; 3 ♂, Tsagaankhayrkhan Soum, Onchun Gol, 15 km ESE Tsagaankhayrkhan, N49.38851, E94.44290, 1832 m, 20–21.vii.2010, MAIST, MAIS 2010072001. Zavkhan Aimag: 9 ♂, Ikh-Uul Soum, Ideriin Gol ~ 28 km E Tosontsengel, N48.71968, E98.65184, 1654 m, 19–20.vii.2004, SRPT, SRP 04071903; 4 ♂, Tosontsengel Soum, Delgarakhiin Gol ~ 16 km S Tosontsengel, N48.61518, E98.23096, 1761 m, 20–21.vii.2004, SRPT, SRP 04072001; 17 ♂, Tosontsengel Soum, Gunza Gol ~ 38 km SW Tosontsengel, N48.62624, E97.89617, 1831 m, 21–22.vii.2004, SRPT, SRP 04072102; 1 ♂, 3 ♀, Telmen Soum, Ideriin Gol ~ 15 km SSW Telmen/Ovogdii, N48.53255, E97.52093, 1823 m, 24–25.vii.2004, SRPT, SRP 04072402; 1 ♂, 2 ♀, Ider Soum Ideriin Gol ~ 4 km NE Zuunmod/Ider, N48.24995, E97.40627, 1929 m, 22–23.vii.2004, SRPT, SRP 04072203; 2 ♂, 1 ♀, Ider Soum, Dogshin/Nogoon Nuur~ 21km SE Zuunmod/Ider, N48.06257, E97.55064, 2054 m, 23–24.vii.2004, SRPT, SRP 04072302; 2 ♂, Ider Soum, ponds by Ideriin Gol at Darkhjan Uul Brigade ~ 12 km SE Zuunmod/Ider, N48.13246, E97.48425, 2025 m, 23.vii.2004, SRPT, SRP 04072301; 1 ♂, 3 ♀, Ikh-Uul Soum, Deed Tsetsuukhiin Gol ~ 10 km NW Solongotiin Davaa, N48.36436, E98.92737, 2072 m, 25.vii.2004, SRPT, SRP 04072502.

Elevation range in Mongolia. Adults were collected at altitudes ranging from 600 m to 2800 m.

Period of activity. Adults are active from the mid-June through to early August.

Known distribution. China, Kazakhstan, Mongolia ( Map 14 ) and Russia (western and eastern parts of Siberia).

Male. Body length 13.6–16.1 mm, wing length 14.6–17.3 mm. General body coloration brownish yellow.

Head. Gray. Vertex and occiput light gray with narrow median line. Rostrum coloration varies from yellow to brownish yellow, nasus conspicuous. Antenna 13-segmented, if bent backward reaching beyond base of wing. Scape and pedicel yellow, flagellomeres 1–3 bicolorous, succeeding dark brown. Each flagellomere, except first one, with weak basal enlargement and deeply incised. Apical flagellomere very small, reduced, distinctly shorter than preceding flagellomere. Verticils as long as respective segments. Palpus with 2 basal segments brownish, succeeding dark brown.

Thorax. Brownish gray. Pronotum light brown with broad brown median line. Prescutum and presutural scutum gray with 4 brownish longitudinal stripes indistinctly bordered by dark brown. Intermediate pair separated. Postsutural scutum gray, scutal lobe with 2 spots indistinctly bordered by brown. Scutellum light brown with brown median line. Postnotum gray pruinose with brown median line. Pleura brownish gray pruinose. Coxae yellowish, dusted with gray. Trochanters yellowish. Femora yellowish with narrowly darkened distal part. Tibiae yellowish with darkened distal part. Tarsal segments brown. Tarsal claws without tooth. Wing patterned with brown. Halter with pale stem and brown knob.

Abdomen. Yellow, with conspicuous dark brown dorsal and lateral stripes. Segment 1 sparsely dusted with gray. Lateral and caudal borders of tergites pale. Sternites sparsely dusted.

Hypopygium. Yellow. Tergite 9 forming large flattened plate ( Fig. 174 ). Posterior margin of tergal plate with broad U-shaped blackened notch, lateral lobes obtusely rounded. Dorsal surface of tergal plate with 2 oblique carina, converging posteriorly. Gonocoxite shaped as large flattish plate with apex obtuse blackened, inconspicuous lateral tooth in middle of posterior surface ( Fig. 175 ). Outer gonostylus widest at middle with apex rounded ( Fig. 176 ). Inner gonostylus large, mid-dorsal edge with indistinct pale plate, upper beak sclerotised and rounded, lower beak sclerotised and swollen, both beaks are separated by deep notch ( Fig. 177 ). Gonocoxal fragment ( Fig. 178 ) with medial sclerites flattened posteriorly, anteriorly fused into weak base and together forming V-shaped structure. Lateral sclerite very small and inconspicuous. Aedeagal guide in shape of triangular broad-based groove, gonapophyses short and wrinkled, adminicular rods broad ( Fig. 179 ). Sperm pump with central vesicle flattened ( Fig. 180 ). Compressor apodeme shaped as large plate without median incision. Posterior immovable apodeme about same length as compressor apodeme with rounded apex. Anterior immovable apodeme rounded. Aedeagus shaped as elongate tube, about 7.4 times as long as sperm pump, basally and medially dark brown, apically passing into yellow ( Figs 12 , 181 ). Distal part with deep lateral and ventral incisions, appearing trident-shaped ( Fig. 181 ).

Female. Body length 21.2–27.5 mm, wing length 2.2–2.7 mm. Female differs from male by reduced wings and elongate abdomen. Antenna short, if bent backward reaching base of wing. Antenna brown, flagellomeres except first one with small weak basal enlargements. Dorsal abdominal stripe broad, black.

Ovipositor ( Figs 182–184 ). Tergite 10 shiny brown. Cercus brown, longer than tergite 10, broad at base with tip acute, outer margin with indistinct serration ( Fig. 182 ). Hypovalva short, sclerotized stout projection ( Fig. 183 ). Sternite 9 with anterior part distinctly broadened, posterior part long and acute. Furca anteriorly dark brown and narrow, posteriorly membranous, except median yellowish-brown area medially. Bursa copulatrix and spermathecal ducts without sclerotisation ( Fig. 184 ). Spermatheca spherical ( Fig. 15 ).

Remarks. The nominative subspecies T. (V.) virgatula virgatula is widely distributed in Mongolia, also known from Russia (west and east of Siberia), Kazakhstan and China (northwest) ( Oosterbroek 2019). The subspecies T. (V.) virgatula montivaga was described by Savchenko (1960) from Alatau mountain range in Kazakhstan and also known from Kyrgyzstan and China (Xinjiang province). According to Savchenko (1960, 1964) the subspecies differs by darker body coloration, males with longer antennae reaching the base of the abdomen; flagellomeres with basal enlargement smaller than in T. (V.) virgatula virgatula with shallower incisions; tergite 9 with oblique carina much darker ( Savchenko 1964, fig. 151–2); inner gonostylus with beak massive ( Savchenko 1964, fig. 152–2). These characters clearly indicate that T. (V.) virgatula montivaga should be elevated to full species level, but study of the male and female terminalia, including type specimens are needed to establish its taxonomic treatment.

Figures

FIGURES 10–13. Aedeagus examples of Mongolian T. (Vestiplex), lateral view. 10. T. (Vestiplex) scripta; 11. T. (Vestiplex) mediovittata; 12. T. (Vestiplex) virgatula; 13. T. (Vestiplex) kiritshenkoi.

FIGURES 14–27. Spermathecae of Mongolian T. (Vestiplex). 14. T. (Vestiplex) longitudinalis; 15. T. (Vestiplex) virgatula; 16. T. (Vestiplex) pallitergata; 17. T. (Vestiplex) laccata; 18. T. (Vestiplex) balioptera; 19. T. (Vestiplex) nubeculosa; 20. T. (Vestiplex) scripta; 21. T. (Vestiplex) kiritshenkoi; 22. T. (Vestiplex) mediovittata; 23. T. (Vestiplex) subcentralis; 24. T. (Vestiplex) sintenisi; 25. T. (Vestiplex) coronifera; 26. T. (Vestiplex) tchukchi; 27. T. (Vestiplex) leucoprocta.

FIGURES 174–181. Hypopygium of male T. (Vestiplex) virgatula. 174. Tergite 9, dorsal view; 175. Gonocoxite, lateral view; 176. Left outer gonostylus, lateral view; 177. Left inner gonostylus, lateral view; 178. Gonocoxal fragment, dorsal view; 179. Aedeagal guide, lateral view; 180. Sperm pump, dorsal view; 181. Distal part of aedeagus, lateral view. Abbreviation: gon, gonapophysis. Scale of 174–180 = 0.5 mm, 181 = 0.25 mm.

FIGURES 182–184. Ovipositor of female T. (Vestiplex) virgatula. 182. Ovipositor, left lateral view; 183. Sternite 8 with hypovalvae, ventral view; 184. Sternite 9, furca and bursa copulatrix, dorsal view. Scale of 182 = 0.8 mm, 183–184 = 0.5 mm.

MAPS 13–14. Distribution maps of T. (Vestiplex) crane flies in Mongolia. 13. T. (Vestiplex) longitudinalis; 14. T. (Vestiplex) virgatula.