Enicospilus antefurcalis (Szepligeti, 1908)

Shimizu, So, 2017, Description of a new species and revised key to species of the Enicospilus antefurcalis species-group from Japan (Hymenoptera: Ichneumonidae: Ophioninae), Acta Entomologica Musei Nationalis Pragae 57 (1), pp. 183-194 : 191-192

Key to species of the Enicospilus antefurcalis species-group

1 Central sclerite completely absent. Oceanic. .......... E. kalveus Gauld & Mitchell, 1981

– Central sclerite present ( Fig. 5 ). ..................................................................................... 2

2 Proximal corner adjacent to Rs+2r of marginal cell with wide glabrous area. Central sclerite elongate linear-oval. Eastern Palaearctic and Oriental. ........................................ .............................................................................................. E. sauteri (Enderlein, 1921)

– Proximal corner of marginal cell entirely setose, without glabrous area. Central sclerite various, oval to circular ( Fig. 5 ). .................................................................................... 3

3 Proximal and distal sclerites separated, not confluent. ................................................... 4

– Proximal and distal sclerites confluent ( Fig. 5 ). ........................................................... 10

4 Lower face 0.9 times as wide as high. Clypeus wide and 1.9 times as wide as high. AI = 0.3. Australasian. .............................................. E. ypsilon Gauld & Mitchell, 1981

– Lower face 0.7–0.8 times as wide as high. Clypeus normal and 1.5–1.8 times as wide as high. AI = 0.4–0.5. .......................................................................................................... 5

5 ICI = 0.8. Distal sclerite weak. Fenestra rather small. Australasian. ................................ ................................................................................... E. xuthus Gauld & Mitchell, 1981

– ICI = 0.4–0.6. Distal sclerite strong. Fenestra moderately large. ................................... 6

6 Central sclerite large, about 40% as wide as proximal sclerite and its minimum diameter larger than minimum distance between it and Rs+2r. Afrotropical and Oriental. .......... 7

– Central sclerite of moderate size, less than 20% as wide as proximal sclerite and its minimum diameter smaller than minimum distance between it and Rs+2r. ...................... 8

7 Central sclerite weakly sclerotized and pigmented. Upper tooth of mandible 2.1–2.3 times as long as lower tooth. ...................................... E. aciculatus (Taschenberg, 1875)

– Central sclerite moderately to strongly sclerotized and strongly pigmented. Upper tooth of mandible 1.3–1.4 times as long as lower tooth. ......... E. laqueatus (Enderlein, 1921)

8 Posterior part of metasoma black. Distal 0.8 of outer surface of fore tibia with moderately dense spines. Afrotropical. .................................................. E. watshami Gauld, 1982

– Posterior part of metasoma not black. Distal half of outer surface of fore tibia with sparse spines. ............................................................................................................................. 9

9 Central sclerite oval, its longer axis parallel to Rs+2r. Face yellowish-white except for central area. Mandibles with sparse setae. Afrotropical. ........... E. polemus Gauld, 1982

– Central sclerite oval to suboval, its longer axis at right angle to Rs+2r. Face entirely orange-yellow. Mandibles with dense setae. Afrotropical and Oceanic. ........................... .................................................................................... E. ruscus Gauld & Mitchell, 1978

10 Central sclerite positioned in middle of fenestra. Corner formed by distal margin of proximal screrite and proximal part of distal sclerite, more or less angulate. Australasian. .................................................................................... E. xaivus Gauld & Mitchell, 1981

– Central sclerite positioned in distal part of fenestra ( Fig. 5 ). Corner formed by distal margin of proximal sclerite and proximal part of distal sclerite, evenly curved ( Fig. 5 ). ...................................................................................................................................... 11

11 Mesosoma , coxae, T1, T2, T5–8 dark-brown to black ( Fig. 5 ). Metapleuron roughly punctostriate ( Fig. 6 ). Eastern Palaearctic. ....................................... E. kikuchii sp. nov.

– Body entirely yellowish-brown or posterior part of metasoma blackish. Metapleuron uniformly punctate or finely punctostriate. ....................................................................... 12

12 AI = 0.2–0.3, CI = 0.2–0.3. Distal sclerite weak. Central sclerite oval and large, its longer axis reclivous and parallel to M between 2m-cu and 3rs-m. Oriental. .............................. ..................................................................................... E. marathwadensis Nikam, 1980

– AI = 0.3–1.1, CI = 0.3–0.6. Distal sclerite more or less strong. Central sclerite circular to oval, if oval more or less small and its longer axis inclivous, not parallel to M between 2m-cu and 3rs-m. .......................................................................................................... 13

13 Mandible twisted 10–20°. Metapleuron punctostriate or punctate. Posterior part of metasoma usually black. Australasian, Eastern Palaearctic, Oceanic, and Oriental. ................ ..................................................................................... E. melanocarpus Cameron, 1905

– Mandible twisted 25°. Metapleuron closely punctate. Posterior part of metasoma black or yellowish brown. Afrotropical. ................................................................................ 14

14 Posterior part of metasoma black. Central sclerite small, its maximum diameter smaller than minimum distance between it and Rs+2r. AI = 0.5–1.0. ........................................... .......................................................................................... E. bicoloratus Cameron, 1912

– Posterior part of metasoma yellowish-brown. Central sclerite moderately large, its maximum diameter larger than minimum distance between it and Rs+2r. AI = 0.3–0.5. ......... ..................................................................................... E. antefurcalis (Szépligeti, 1908)

Figures

Figs 2–5. Partial views of Enicospilus kikuchii sp. nov. (holotype). 2–4 – head in frontal (2), lateral (3), and dorsal views (4); 5 – part of fore wing.

Figs 6–11. SEM images of Enicospilus kikuchii sp. nov. (holotype). 6 – mesosoma in lateral view; 7 – propodeum in dorso-lateral view; 8 – fore tibia in lateral view; 9 – anterior part of T2 in lateral view.