Glyptothorax fuscus Fowler 1934

Ng, Heok Hee & Kottelat, Maurice, 2016, The Glyptothorax of Sundaland: a revisionary study (Teleostei: Sisoridae), Zootaxa 4188 (1), pp. 1-92 : 24-32

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Glyptothorax fuscus Fowler 1934


Glyptothorax fuscus Fowler 1934 View in CoL

( Figs. 12 View FIGURE 12 , 16 View FIGURE 16 )

Glyptothorax fuscus Fowler, 1934: 89 View in CoL , Figs. 31–33 View FIGURE 31 View FIGURE 32 View FIGURE 33 (type locality: Chantaburi, Thailand); Smith, 1945: 403; Suvatti, 1981: 95; Jiang et al., 2011: 280; Rainboth et al., 2012: 70.

Glyptosternum telchitta View in CoL (non Hamilton, 1822)— Duncker, 1904: 174.

Glyptothorax platypogon View in CoL (non Valenciennes, 1840)—Herre & Myers, 1937: 68; Hora & Gupta, 1941: 35.

Glyptothorax telchitta View in CoL (non Hamilton, 1822)— Fowler, 1938: 48.

Glyptothorax laosensis View in CoL (non Fowler, 1934)— Fowler, 1939: 57.

Glyptothorax majus (non Boulenger, 1894)— Herre, 1940: 35; Tweedie, 1940: 70; Hora & Gupta, 1941: 34.

Glyptothorax callopterus Smith, 1945: 400 View in CoL , Fig. 87 (type locality: waterfall stream on Khao Chong, near Trang, Thailand); Suvatti, 1981: 95 (in part); Cramphorn, 1983: 20; Jiang et al., 2011: 280; Amirrudin & Zakaria-Ismail, 2014: 414.

Glyptothorax major View in CoL (non Boulenger, 1894)— Smith, 1945: 401, Fig. 88; Suvatti, 1950: 303; Alfred, 1961: 6, Pl. 2 Fig. 6 View FIGURE 6 ; 1966: 39, Pl. 6 Fig. 3 View FIGURE 3 ; Bishop, 1973: 350; Mohsin et al., 1977: 76; Mizuno & Furtado, 1982: 323; Mohsin & Ambak, 1982: 102, Fig. 2 View FIGURE 2 ; 1983: 123, Fig. 90; Zakaria-Ismail, 1984: 25; 1993: 208, Pl. 6a; Roberts, 1989: 134 (in part); Lim et al., 1990: 47; Zakaria-Ismail & Lim, 1995: 323, Pl. 4d; Sim, 2002: 61; Ambak et al., 2010: 86.

Glyptothorax prashadi View in CoL (non Mukerji, 1932)— Smith, 1945: 402; Menon 1955: 42 (in part).

Glyptothorax aff. major View in CoL —Ng & Tan, 1999: 362.

Glyptothorax cf. major View in CoL — Kottelat et al., 1992: 10; Kano et al., 2013: 179.

Glyptothorax sp. cf. major —Miyazaki et al., 2013: 1041, Fig. 3 View FIGURE 3 E.

Material examined. SOUTHEAST THAILAND: ANSP 59358 View Materials (holotype), 59.7 mm SL; Chantaburi . UMMZ 246371 (23), 31.9–63.1 mm SL; Chantaburi: Amphoe Soi Dau, Tambon Pathong , Klong Thepprathan. ZRC 35788–35790 View Materials (3), 21.5–33.3 mm SL ; CMK 10663 (4), 23.4–38.0 mm SL; Chantaburi: about 1 km on road to Namtok Phliu after Chantaburi Trat highway. ZRC 35757 (1), 39.7 mm SL ; CMK 10721 (1), 39.7 mm SL; Trat: Klong Fit at Ban Kraduk Chang , Trat –Bo Rai road about 1–2 km after junction with road 3271.

MALAY PENINSULA: USNM 109820 (holotype of G. callopterus ) 51.8 mm SL; Thailand: Trang, waterfall stream at Khao Chong. USNM 109819 View Materials (2 paratypes of G. callopterus ), 48.0– 48.2 mm SL ; Thailand: Trang, mountain stream at Khao Chong. UF 183902 (1), 47.3 mm SL ; Thailand: Phangnga, Bangdook River , off road around PHG 3059, 8 °35'58"N 98°26'36"E. UF 183901 (4), 39.4–49.0 mm SL ; Thailand: Phangnga, Bangdook River , small creek off side road around PHG 3059, 8 °34'28"N 98°26'40"E. CMK 22145 (4), 22.0– 45.8 mm SL ; Thailand: Phangnga, Khlong Bok Pui, Ban Bok Pui at junction of roads 4090 and 4240, 8°34'25"N 98°34'53"E. UF 183904 (1), 50.9 mm SL GoogleMaps ; Thailand: Phangnga, Phangnga River, S of road 4090 behind small house, 8°32'20"N 98°29'14"E. UMMZ 195916 View Materials (1), 37.6 mm SL GoogleMaps ; Thailand: Phangnga, tributary of Khlong Ha Ngam , 1 km S of Ban Ang. UF 183324 (5), 33.6–59.4 mm SL ; Thailand: Nakhon Si Thammarat, Ta Pae Kao river at Chang Klang, off road 4015, Tapi River drainage, 8°20'56"N 99°41'32"E. ANSP 59373 View Materials (1), 42.6 mm SL GoogleMaps ; Thailand: Nakhon Si Thammarat. ANSP 76834 View Materials (48), 33.2–58.9 mm SL ; ANSP 76835 (3), 28.8–36.7 mm SL; Thailand: Trang, Waterfall at Khao Chong , 19 km E of Thap Thiang. ZRC 42162 (3), 29.2–37.5 mm SL ; CMK 12117 (3), 24.9-32.2 mm SL; Thailand: Satun, stream in Ban Kong Kruat. ZRC 42098 (2), 25.9–28.5 mm SL ; CMK 12045 (11), 34.1–43.4 mm SL; Thailand: Narathiwat, stream along road branching W at about 7 km on road from Waeng to Ban Bu Ke Ta. ZRC 42123 (14), 21.5–52.5 mm SL ; CMK 12077 (14), 15.1–48.8 mm SL; Thailand: Narathiwat, Mae Nam Sungai Kolok at Ban Sae , 7 km S of Ban Bu Ke Ta. ZRC 41063 (24), 10.1–58.9 mm SL ; Perak: Sungai Kenderong at Gerik , 5°25'29"N 101°7'39"E. ZRC 23351–23354 View Materials (4), 44.6–48.9 mm SL GoogleMaps ; ZRC 38732 (82), 18.3–56.7 mm SL; ZRC 41877 (40), 14.0–56.0 mm SL; CMK 8180 (34), 29.8–52.0 mm SL; Malaysia: Terengganu, Sungai Brang , outside Sekayu Waterfall Park. CAS-SU 31004 View Materials (1), 52.0 mm SL ; Malaysia: Perak, Plus River near Pos Legap. ZRC 41042 (38), 11.9–26.0 mm SL ; Malaysia: Perak, Sungai Korbu at Jalong , 4°50'58"N 101°11'8"E. ZRC 39973 (10), 15.4–35.6 mm SL GoogleMaps ; Malaysia: Perak, tributary of Sungai Korbu in the vicinity of Jalong. USNM 229305 View Materials (1), 49.6 mm SL ; Malaysia: Terengganu, forest streams near Kampung Ayer Puteh (29–35 km inland from Chukai). CAS-SU 39344 View Materials (41), 27.6–53.6 mm SL ; Malaysia: Perak, Tapah. CAS-SU 39343 View Materials (3), 17.9–55.4 mm SL ; Malaysia: Perak, Batang Padang. ZRC 41225 (2), 25.5–27.2 mm SL ; Malaysia: Pahang, Sungai Kla at Raub. CAS- SU 32677 (1), 49.0 mm SL ; Malaysia: Selangor, W of Genting Sempah , 29 km E of Kuala Lumpur. CAS-SU 32678 View Materials (1), 48.2 mm SL ; Malaysia: Pahang, Benus River , E of Genting Sempah. ZRC 42848 (2), 25.6–44.1 mm SL ; Johor: near Mersing. ZRC 42757 (4), 27.6–52.1 mm SL ; Johor: Sungai Kahang and tributary ca. 44.4 km from Mersing turnoff to Kluang just before side road to Endau Rompin Taman Negara , km 96 from Mersing to Batu Pahat , 2°3'56"N 103°31'35"E. ZRC 40257 (2), 41.6–44.5 mm SL GoogleMaps ; Malaysia: Johor, tributary of Sungai Batang Pinang, 272 km towards Kluang. CAS-SU 3 View Materials 1003 (1), 41.9 mm SL ; Malaysia: Johor, Muar. CAS-SU 39342 View Materials (4), 36.7–40.6 mm SL ; Malaysia: Johor, Simpang Renggam. USNM 229263 View Materials (1), 49.2 mm SL ; Malaysia: Johor, small forest tributaries of Sungai Sedili Besar , 10–22 km N of Kota Tingi (on road to Mersing). CAS-SU 32673 View Materials (2), 22.4–30.0 mm SL ; Malaysia: Johor, Mawai district . FMNH 62019 View Materials (1), 20.2 mm SL ; FMNH 62020 (1), 17.6 mm SL; Malaysia: Johor, Mawai Estate. FMNH 62317 View Materials (6), 15.3–25.3 mm SL ; Malaysia: Johor, small stream along road from Kota Tinggi to Mawai. CAS-SU 132675 View Materials (3), 40.4–54.4 mm SL ; Malaysia: Johor, 8 km N of Kota Tinggi. ZRC 39951 (2), 40.4–48.5 mm SL ; Malaysia: Johor, Kota Tinggi area , Sungai Tementang.

SINGAPORE: CAS-SU 32674 View Materials (1), 38.9 mm SL; Mandai Road.

BANGKA: ZRC 30736–30742 View Materials (7), 34.6–45.0 mm SL; CMK 9619 (9), 28.5–45.6 mm SL; stream 9 km E of Mentok . ZRC 31263–31264 View Materials (2), 24.6–48.7 mm SL; CMK 9646 (1), 44.9 mm SL; 28 km N of Payung .

SUMATRA: UF 166633 (1), 29.3 mm SL ; Sumatera Selatan: Air Rambang , 3°33'2"S 104°15'50"E. UF 166632 (1), 34.6 mm SL GoogleMaps ; Sumatera Selatan: Air Kikim Kecil , 3°45'50"S 103°27'34"E. ZRC 53528 (1), 43.3 mm SL GoogleMaps ; Sumatera Selatan: market in Lahat, Lematang (Musi) drainage, 3°48'S 103°32'E. UF 166631 (1), 46.0 mm SL GoogleMaps ; Sumatera Selatan: Air Enai , 4°3'39"S 104°2'52"E. UF 167899 (31), 23.9–58.6 mm SL GoogleMaps ; Lampung: Way Besai , 4°45'27"S 104°33'20"E. UF 166638 (1), 46.8 mm SL GoogleMaps ; Lampung: Way Sebetih , 4°37'36"S 104°59'53"E. UF 161617 (2), 51.2–60.3 mm SL GoogleMaps ; Lampung: Way Seputih draiange, 5°4'189"S 104°53'3"E.

Diagnosis. Glyptothorax fuscus differs from all Sundaic congeners except G. prashadi and G. s t i b a ro s in having a straight (vs. concave) dorsal margin of the dorsal fin. It is distinguished from G. prashadi in having an acutely (vs. broadly) rounded snout when viewed dorsally ( Fig. 13 View FIGURE 13 ), a more tapering body (caudal peduncle depth 2.3–2.8 times in body depth at dorsal origin vs. 1.8–1.9), expanded distal tips of neural spines forming (vs. not forming) a series of low bumps anterior to adipose fin and a caudal fin with acutely rounded (vs pointed) lobes and hyaline areas in the median fin rays and tips of fin lobes (vs. hyaline areas absent and caudal fin with same color pattern throughout; Fig. 14 View FIGURE 14 ), and from G. stibaros in having a smooth (vs. serrated) posterior margin of the dorsalfin spine, smaller tubercles in the abdominal region ( Fig. 15 View FIGURE 15 ) and a less tapering body, as manifested by the smaller ratio between the body and caudal peduncle depths (caudal peduncle depth 2.3–2.8 times in body depth at dorsalfin origin vs. 3.4–3.8). The following unique combination of characters further distinguishes G. f u s c u s from Sundaic congeners: head width 18.3–23.3% SL; anteromedial striae in thoracic adhesive apparatus absent; width of thoracic adhesive apparatus 1.3–1.6 times in its length; dorsal-to-adipose distance 14.8–20.6% SL; body depth at anus 15.6–20.7% SL; straight dorsoposterior margin of adipose fin; post-adipose distance 15.7–20.8% SL; caudal peduncle depth 7.4–10.9% SL; caudal-fin length 28.7–33.9% SL; dark bands on caudal-fin lobes running divergent to longitudinal axis of lobes; lacking both prominent pale midlateral stripe on body and dark vertical bars at adipose-fin base and base of caudal fin.

Description. Morphometric data in Table 5 View TABLE 5 . Head depressed; body subcylindrical. Dorsal profile rising steeply from tip of snout to origin of dorsal fin, then sloping gently ventrally from origin of dorsal fin to end of caudal peduncle. Ventral profile straight to anal-fin base, then sloping gently dorsally from anal-fin base to end of caudal peduncle. Anus and urogenital openings located at vertical through middle of adpressed pelvic fin. Skin tuberculate, with tubercles of on sides of body generally larger than those on dorsal surfaces of head and body. Lateral line complete and midlateral. Vertebrae 16+17=33 (4), 17+16=33 (2), 16+18=34 (6), 17+17=34 (17), 18+16=34 (2), 17+18=35 (8) or 18+17=35 (3).

Head depressed and broad, triangular when viewed laterally. Snout prominent. Anterior and posterior nares large and separated only by base of nasal barbel. Gill openings broad, extending from ventral margin of posttemporal to isthmus. First branchial arch with 2+6 (7), 2+7 (12) or 3+8 (1) rakers. Bony elements of dorsal surface of head covered with thick, tuberculate skin. Eye ovoid, horizontal axis longest; located entirely in dorsal half of head.

Barbels in four pairs. Maxillary barbel long and slender, extending to middle of pectoral-fin base. Nasal barbel slender, extending to midway between its base and anterior orbital margin. Inner mandibular-barbel extending to midway between its base and that of pectoral spine. Outer mandibular barbel extending to two-thirds of distance between its base and that of pectoral spine.

Mouth inferior, premaxillary tooth band partially exposed when mouth is closed. Oral teeth small and villiform, in irregular rows on all tooth-bearing surfaces. Premaxillary teeth appearing in single broad semilunate band. Dentary teeth in a single crescentic band, consisting of two separate halves tightly bound at midline.

Thoracic adhesive apparatus consisting of keratinized striae in an ovate field extending from isthmus to just beyond posterior limit of pectoral-fin base ( Fig. 3 View FIGURE 3 e). Narrow, chevron-shaped medial pit on posterior third. Striae around posterior half of median depression anastomosing.

Dorsal fin located above anterior third of body, with I,6 (20) rays; fin margin straight; spine short and straight, smooth on anterior and posterior margins. Adipose fin with anterior margin straight or slightly concave and posterior margin straight. Caudal fin strongly forked, with slightly rounded tips, lower lobe slightly longer than upper lobe and i,7,8,i (20) principal rays. Procurrent rays symmetrical and extending only slightly anterior to fin base. Anal-fin base vertically opposite adipose-fin base. Anal fin with straight anterior margin and straight or slightly concave posterior margin; with iv,8 (1), iv,8,i (3), iv,9 (4), v,8,i (2), iv,9,i (4), iv,10 (2), v,9 (1), iv,10,1 (1), iv, 11 (1) or v,9,i (1) rays. Pelvic-fin origin at vertical through posterior limit of dorsal-fin base. Pelvic fin with slightly convex margin and i,5 (20) rays; tip of adpressed fin not reaching anal-fin origin. Pectoral fin with I,7 (1), I,7,i (16), I,8 (1) or I,8,i (2) rays; posterior fin margin slightly concave; anterior spine margin smooth, posterior margin with 3–10 serrations.

Coloration. In 70% ethanol: dorsal and lateral surfaces of head and body reddish brown to dark gray, fading to paler color (beige to light gray) on ventral surfaces. Dorsal and lateral surfaces of head and body with small irregular darker spots in some individuals, particularly towards posterior half of fish ( Fig. 16 View FIGURE 16 ). A thin, pale (usually cream) mid-dorsal stripe extending from base of last dorsal-fin ray to origin of adipose fin; stripe indistinct or nearly absent in some individuals. Laterosensory pores rimmed in beige, imparting appearance of a diffuse light brown midlateral line in some individuals. Dorsal and ventral surfaces of caudal peduncle slightly paler in some individuals. All fins with rays proximally brown or gray, becoming hyaline more distally, and diffuse melanophores on fin membranes. Pectoral and pelvic fins with brown or gray on base of fin rays and hyaline posterior margin. Anal fin with brown base; brown spot on anterior third of fin present in some individuals. Adipose fin brown with hyaline distal margin. Base of caudal fin with dark crescent in most individuals. Each caudal-fin lobe with irregular, elongate brown or gray blotch on about half of outer fin rays; most of inner rays of lobes hyaline. Maxillary and nasal barbels brown to gray dorsally, beige ventrally. Mandibular barbels beige.

Distribution. Glyptothorax fuscus is found in forested hillstreams throughout southern Sumatra (from the Musi River drainage southwards to the short coastal drainages that drain the southern tip of the island), Bangka, the Malay Peninsula (from the southern tip of the peninsula northwards to the Tapi River drainage on the eastern side and to Phangnga on the western side), as well as the short coastal rivers draining the southeastern face of the Khao Banthat (Cardamom) Mountains in southeastern Thailand ( Fig. 17). This species was also found in Singapore, but has since been extirpated there (Ng & Lim, 1997).

Habitat and biology. Glyptothorax fuscus is typically found in lower elevation hillstreams and streams draining the foothills throughout its range. Such streams typically have a moderate to swift current, a sandy bottom and thick vegetation on the banks. This species feeds on aquatic invertebrates (Kano et al., 2013).

Comparisons. Besides G. prashadi (which has already been compared with G. f u s cu s in the diagnosis), there are seven congeners known from the Malay Peninsula and Sumatra: G. amnestus , G. famelicus , G. k e l u k, G. ketambe , G. platypogonides , G. plectilis and G. s c h m i d t i. Glyptothorax fuscus further differs from G. amnestus in having a more tapering body as manifested by the larger ratio between the body and caudal peduncle depths (caudal peduncle depth 2.3–2.8 times in body depth at dorsal-fin origin vs. 1.4–2.1), from G. famelicus , G. ke t a m b e and G. schmidti in lacking (vs. with) a prominent pale midlateral stripe on the body, and in having a deeper body (depth at anus 15.6–20.7% SL vs. 11.4–15.7); it further differs from G. famelicus in having a deeper caudal peduncle (7.4–10.9% SL vs. 5.8–7.3). Glyptothorax fuscus is further distinguished from G. keluk in having a deeper body (depth at anus 15.6–20.7% SL vs. 14.0–15.8) and a straight (vs. convex) dorsoposterior margin of the adipose fin, from G. platypogonides in having a deeper body (depth at anus 15.6–20.7% SL vs. 12.1–14.8), and from G. plectilis in lacking (vs. having) anteromedial striae in the thoracic adhesive apparatus. Among congeners known only from Borneo, it further differs from G. decussatus in lacking (vs. having) anteromedial striae in the thoracic adhesive apparatus, having a more tapering body (caudal peduncle depth 2.3–2.8 times in body depth at dorsal-fin origin vs. 2.0), from G. exodon in having the premaxillary toothband not more than three-quarters (vs. almost entirely) exposed when the mouth is closed, a wider head (18.3–23.3% SL vs. 16.0–18.1), a shorter dorsal-toadipose distance (14.8–20.6% SL vs. 21.9–26.3), a shorter post-adipose distance (15.7–20.8% SL vs. 21.5–24.0) and a deeper body (depth at anus 15.6–20.7% SL vs. 13.8–15.8), and from G. major in lacking (vs. having) anteromedial striae in the thoracic adhesive apparatus. Glyptothorax fuscus is further distinguished from G. nieuwenhuisi in having a shorter, wider thoracic adhesive apparatus (width 1.3–1.6 times in its length vs. 1.6–1.9), more tapering body (caudal peduncle depth 2.3–2.8 times in body depth at dorsal-fin origin vs. 1.7–2.4), and from G. p i c t u s in lacking (vs. having) dark vertical bars at adipose-fin base and base of caudal fin. Among congeners known only from Java, G. f u s c u s further differs from G. platypogon in having a straight (vs. convex) dorsoposterior margin of the adipose fin, a longer caudal fin (28.7–33.9% SL vs. 20.1–27.0), and from G. robustus in having (vs. lacking) a medial pit in the thoracic adhesive apparatus, broader skin ridges on the thoracic adhesive apparatus (compare Figs. 3 View FIGURE 3 e and 3o) and lacking (vs. having) a prominent dark longitudinal stripe running through each caudal-fin lobe.

Glyptothorax coracinus is a species that is possibly sympatric with G. f u s c u s for part of its range (southeastern Thailand). Glyptothorax fuscus differs from G. coracinus in having fewer total vertebrae (33–35 vs. 36–37) and in having a mottled brown or gray (vs. uniform blackish brown) color pattern.

Remarks. Glyptothorax fuscus is the most widely distributed Sundaic species that has often been misidentified as G. m a j o r, a species known only from Borneo. Kottelat (2001) used this name (G. f u s c u s) for a species from the Laotian Mekong, which we now reidentify as G. l a m p r i s.

TABLE 5. Morphometric data for Glyptothorax fuscus (n = 20).

  Range Mean±SD
Standard length (mm) 42.8–58.9  
Predorsal length 36.7–42.7 39.3±1.53
Preanal length 63.0–68.2 65.7±1.49
Prepelvic length 47.4–54.8 50.9±1.91
Prepectoral length 19.9–26.1 22.6±1.68
Length of dorsal-fin base 12.8–16.0 14.6±0.83
Dorsal-fin spine length 14.6–20.8 17.4±1.70
Length of anal-fin base 15.0–19.6 17.2±1.22
Pelvic-fin length 12.6–17.7 14.5±1.26
Pectoral-fin length 21.2–27.9 24.1±1.43
Pectoral-fin spine length 17.0–22.7 19.3±1.45
Caudal-fin length 28.7–33.9 30.7±1.46
Length of adipose-fin base 13.1–21.0 16.8±1.68
Dorsal to adipose distance 14.8–20.6 17.7±1.74
Post-adipose distance 15.7–20.8 17.9±1.45
Length of caudal peduncle 16.7–22.8 18.8±1.58
Depth of caudal peduncle 7.4–10.9 8.7±0.86
Body depth at anus 15.6–20.7 18.5±1.49
Body depth at dorsal-fin origin 17.4–24.4 21.6±2.07
Head length 26.8–30.6 28.5±1.03
Head width 18.3–23.3 20.8±1.11
Head depth 15.3–19.3 16.8±1.05
Snout length 46–53 49±1.8
Interorbital distance 24–33 29±2.3
Eye diameter 6–13 9±1.6
Nasal barbel length 17–29 22±2.8
Maxillary barbel length 82–109 95±7.2
Inner mandibular barbel length 28–48 40±4.7
Outer mandibular barbel length 42–67 57±6.4

Academy of Natural Sciences of Philadelphia


Zoological Reference Collection, National University of Singapore


Smithsonian Institution, National Museum of Natural History


Peper Harow


University of Michigan, Museum of Zoology


California Academy of Sciences, Stanford University Collection


Field Museum of Natural History














Glyptothorax fuscus Fowler 1934

Ng, Heok Hee & Kottelat, Maurice 2016

Glyptothorax aff. major

Tan 1999: 362

Glyptothorax cf. major

Kottelat 1992: 10

Glyptothorax fuscus

Rainboth 2012: 70
Jiang 2011: 280
Suvatti 1981: 95
Smith 1945: 403

Glyptothorax callopterus

Amirrudin 2014: 414
Jiang 2011: 280
Cramphorn 1983: 20
Suvatti 1981: 95
Smith 1945: 400

Glyptothorax major

Ambak 2010: 86
Sim 2002: 61
Zakaria-Ismail 1995: 323
Zakaria-Ismail 1984: 25
Mizuno 1982: 323
Mohsin 1982: 102
Mohsin 1977: 76
Bishop 1973: 350
Alfred 1961: 6
Suvatti 1950: 303
Smith 1945: 401

Glyptothorax prashadi

Menon 1955: 42
Smith 1945: 402

Glyptothorax platypogon

Hora 1941: 35

Glyptothorax majus

Hora 1941: 34
Herre 1940: 35

Glyptothorax laosensis

Fowler 1939: 57

Glyptothorax telchitta

Fowler 1938: 48

Glyptosternum telchitta

Duncker 1904: 174
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