Etheophanus striatus Broun, 1910

Davis, Steven R., Brav-Cubitt, Talia, Buckley, Thomas R. & Leschen, Richard A. B., 2019, Systematics of the New Zealand Weevil Etheophanus Broun (Curculionidae: Molytinae), Zootaxa 4543 (3), pp. 341-374 : 360-361

Etheophanus striatus Broun, 1910

( Figs 1E , 2E , 5E , 14 , 15 )

Etheophanus striatus Broun, 1910: 69 . Type locality: Waimarino. Etheophanus punctiventris Broun, 1914: 231 . Type locality: Mount Hutt and McClennan's Bush, near Methven. New Synonymy.

Diagnosis. Dorsal surfaces coarsely punctate, alveolate microsculpture indistinct. Pronotal vibrissae usually absent (short and present in some specimens). Procoxa without semi-lamellate process, primary dorsal setae decumbent. Eighth sternite of male with each sclerite bearing ca. 4 setae along apical margins; spiculum relictum broad. Aedeagus with setae present to basal 1/3 of parameroid lobes; median lobe in dorsal view not narrowing to apex and subrounded apically with simple edges, in lateral view broad and not strongly curved ventrally near apex; apico-lateral margin of median lobe bearing 1 seta at each corner. Female tergite VIII apically subrounded; eighth sternite with arms of base strongly arcuate, multisetose to basal 2/3 or 1/2. Gonostylus with 3 apical setae. Spermatheca falciform and recurved without globular base.

Comments. Broun based the description of E. striatus on two specimens, one that we designated as the lectotype to stabilise the epithet, and the other, noted by Broun as a sexual variant, was designated as a paralectotype. While the type locality of E. striatus is in the southern half of the North Island, Broun (1914) later described E. punctiventris from Canterbury. Despite the geographic distance separating the type localities, the specimens are morphologically similar and falling within the geographic variation of the species and therefore conspecific and considered as synonymns. Broun based the description of E. punctiventris on three specimens, one that we designated as the lectotype to stabilise the epithet, and the two remaining were designated as paralectotypes.

Distribution. North Island: WO, TO, RI, GB, HB, WI, WN. South Island: NN, BR, MB, WD, NC, MC, SC, CO, DN.

Type material examined. E. striatus : Lectotype ( BMNH): card-mounted, “LECTO-TYPE [round label with dark blue border] // 3138. [in Broun’s hand] // New Zealand. [red underline] Broun Coll. Brit. Mus. 1922-482. // Waimarino. Jany. 1909. [in Broun’s hand] // Etheophanus striatus . [in Broun’s hand] // LECTOTYPE [printed] ♀ Etheophanus striatus Broun, 1910 det. R. Craw, 1987 [in Craw’s hand on orange dayglow label]” . Paralectotype ( BMNH): 1, card-mounted, “var. [in Broun’s hand] // New Zealand. [red underline] Broun Coll. Brit. Mus. 1922- 482. // Waimarino. Jany. 1909. [in Broun’s hand] // Etheophanus striatus . [in Broun’s hand] // PARALECTOTYPE [printed] Etheophanus striatus Broun, 1910 det. R. Craw, 1987 [in Craw’s hand on dark blue label]”. E. punctiventris : Lectotype ( BMNH): card-mounted, “LECTO-TYPE [round label with dark blue border] // 3627. [in Broun’s hand] // New Zealand. [red underline] Broun Coll. Brit. Mus. 1922-482. // Mt Hutt. 12.4.1912. [in Broun’s hand] // Etheophanus punctiventris . [in Broun’s hand] // LECTOTYPE [printed] ♂ Etheophanus punctiventris Broun, 1914 det. R. Craw, 1987 [in Craw’s hand on orange dayglow label]” . Paralectotypes ( BMNH): 1, cardmounted, “3627. [in Broun’s hand] // New Zealand. [red underline] Broun Coll. Brit. Mus. 1922-482. // McLennan’s 23.3.1912 [in Broun’s hand] // Etheophanus punctiventris . [in Broun’s hand] // PARALECTOTYPE [printed] ♀ Etheophanus punctiventris Broun, 1914 det. R. Craw, 1987 [in Craw’s hand on dark blue label]”; 1, same but, “Mt Hutt. 23.4.1912 [in Broun’s hand]” .

Dissected material (NZAC). South Island. BR. Kangaroo Ck, Mawhera, SF, 5km SE Ngahere, 27 Jan. 1972, litter, 72/86, Beech Forest Utilization Project (1 male). WD. 9 mi. S. Franz Josef Town , 7 Feb. 1965, N.A. Walker, litter (1 male) . CO. Rock and Pillar Ra, 1066m, 12 Nov. 1969, J. McBurney, 69/212 (1 male) .

Figures

FIGURES 1A–G. Dorsal habiti of Etheophanus spp. A, E. kuscheli sp. n.; B, E. nitidellus; C, E. optandus; D, E. pingius; E, E. striatus.

FIGURES 2A–G. Lateral habiti of Etheophanus spp. A, E. kuscheli sp. n.; B, E. nitidellus; C, E. optandus; D, E. pingius; E, E. striatus.

FIGURES 5A–G. Key characters of Etheophanus spp. A, E. kuscheli sp. n., apical half of median lobe (dorsal view); B, E. nitidellus, apical half of median lobe (dorsal view); C, E. optandus, apical half of median lobe (dorsal view); D, E. pingius, apical half of median lobe (dorsal view); E, E. striatus, apical half of median lobe (dorsal view); F, E. nitidellus (female), internal view of abdominal ventrites showing pair of apodemes on fifth ventrite; G, E. kuscheli, showing semi-lamellate process on procoxa (arrow).

FIGURES 14A–D. Etheophanus striatus, male terminalia. A, aedeagus, dorsal view; B, aedeagus, lateral view; C, tegmen, dorsal view; D, sternites VIII and IX, showing spiculum relictum.

FIGURES 15A–D. Etheophanus striatus, female terminalia. A, sternite VIII; B, tergite VIII; C, gonocoxites; D, spermatheca.