Inadunata Wachsmuth and Springer, 1885

Głuchowski, Edward, 2002, Crinoids from the Famennian of the Holy Cross Mountains, Poland, Acta Palaeontologica Polonica 47 (2), pp. 319-328 : 320-322

Subclass Inadunata Wachsmuth and Springer, 1885 Order Cladida Moore and Laudon, 1943 Family Botryocrinidae Wachsmuth and Springer, 1886 Genus Cosmocrinus Jaekel, 1898

Type species: Cosmocrinus holzapfeli Jaekel, 1898

Cosmocrinus polonicus sp. nov.

Fig. 2O–U .

Cosmocrinus distinctus Stukalina, 1977 ; Głuchowski 1981a: 49, pl. 3: 12. Cosmocrinus distinctus Stukalina, 1977 ; Głuchowski 1981b: 89–110.

Holotype: GIUS−4−91/3, Fig. 2S .

Type locality: Kadzielnia quarry, Kielce, Holy Cross Mountains.

Type horizon: Marly limestones, Palmatolepis rhomboidea to Palmatolepis marginifera zones.

Derivation of the name: From Latin polonicus, Polish.

Diagnosis.— Cosmocrinus species with ornamented xenomorphic and non−cirriferous stem.

Material.—235 columnals and pluricolumnals.

Dimension of the holotype (in mm).—Columnal diameter (KD) 6.6; luminal diameter (LD) 0.9; crenularial diameter (CD) 5.7; columnal height (KH) 2.2.

Description.—The stem is medium−sized (diameter 2.9– 7.0 mm), xenomorphic and lacking cirri. Three sorts of columnals forming the stem are differentiated in height, shape and ornamentation. Lateral side of the lowest, flat columnals (proxistele?) is covered with distinct, numerous outgrowths or tubercles ( Fig. 2Q, R ). Relatively higher and barrel−shaped columnals (mesistele?) have on their lateral side numerous ridges ( Fig. 2T, U ), arranged more or less regularly. However, lateral side of cylinder−like columnals (dististele?) is covered with more delicate, sometimes bifurcated, ridges ( Fig. 2S ). Articular facet is flat (proxistele) to more or less concave (mesi− to dististele). Small lumen is pentagonal, strongly rounded. Crenularium is composed of rather coarse and long culmina, sometimes dichotomous at the columnal edge ( Fig. 2O, P ).

Remarks.—The species differs from the most similar C. distinctus Stukalina, 1977 from the Late Devonian of Eastern Kasakhstan in lacking cirri. Few specimens from some localities were previously ( Głuchowski 1981a, b) treated as non−cirriferous stem sectors of C. distinctus . However, the new abundant material does not contain any cirriferous nodals and shows that the stalk of the Polish species really lacks cirri.

Occurrence.—This species has been documented from Pa. crepida to S. praesulcata zones: Kadzielnia, Kowala (samples Ko−B and C), Ostrówka, Besówka.

Figures

Fig. 2. A. Kasachstanocrinus sp., GIUS−4−445/4, Kostomłoty, × 20. B, C. Urushicrinus sp., GIUS−4−455/1−2, Psie Górki, sample PG−A, × 15. D. Anthinocrinus sp., GIUS−4−91/33, Kadzielnia, × 12. E. Schyschcatocrinus delicatus Głuchowski, 1993, GIUS−4−455/8, Psie Górki sample PG−A, × 15. F–H. Schyschcatocrinus levis sp. nov., GIUS−4−91/11−13, Kadzielnia; F, G × 15, H (holotype GIUS−4−91/13) × 12. I–J. Stenocrinus altus Sisova, 1988, GIUS−4−91/15−16, Kadzielnia, × 10. K–N. Calleocrinus kielcensis Głuchowski, 1993, GIUS−4−91/45−52, Kadzielnia, × 10. O–W. Cosmocrinus polonicus sp. nov., GIUS−4−91/3−9, Kadzielnia; O–Q × 7, R and S (holotype GIUS−4−91/3) × 5; T, U × 10.