Deltochilum (Aganhyboma) acropyge Bates 1887

Silva, Fernando A. B., Louzada, Júlio & Vaz-De-Mello, Fernando, 2015, A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae), Zootaxa 3925 (4), pp. 451-504 : 471-472

Deltochilum (Aganhyboma) acropyge Bates 1887 new subgeneric placement and new status

( Figs 13 , 46, 91, 106)

Deltochilum acropyge Bates 1887: 36 , plate 2, Fig. 15 (original description) Gillet 1911: 35 (catalogue)

Blackwelder 1944: 202 (checklist)

Blackwelder 1973: 5 (checklist)

Deltochilum (Deltochilum) valgum var. acropyge: Paulian 1938: 269 , 273 (taxonomic comments and identification key) Deltochilum (Deltohyboma) valgum acropyge: Howden 1966: 733 , 735 (identification key, taxonomic comments) Deltochilum valgum acropyge: Howden & Young 1981: 36 , 39 (identification key, description) Cano 1998: 174, 175, 176, 177, Figs 2 and 3 (comment on habitat, distribution and behavior of predation) Halffter 2003: 27, 28, Fig. 2.5 (description)

Description. Body. Head, pronotum, elytra and ventral surface black, with a silky sheen ( Fig. 106 ). Length> 14 mm. Head. Punctures dense, almost touching; foveiform punctures separated at dorsal interocular surface by approximately half diameter. Punctures around clypeus shallower, smaller and more dispersed than those on interocular surface. Lateral margin of gena, nearby eyes, without row of setae or with almost inconspicuous setae. Dorsal interocular distance approximately three times eye width (as in Fig. 61). Thorax. Surface of pronotum with dense foveiform punctures, not touching each other. Punctures on the sides and anterior angles larger and somewhat closer than that of pronotum disc. Discal punctures separated approximately by 1.5 or two times their diameters. Anteromedial punctures smaller and more dispersed than discal punctures. Posteromedial punctures usually separated by at least one diameter. Margin between anterior and lateral angles with row of small setae. Margin between lateral and posterior angles straight, with approximately the same length of margin between anterior and lateral angles. Hypomera with foveiform punctures much more dense on anterior portion; anteromedial punctures separated by distance smaller or equal to one diameter; median portion with sparser punctures. Diagonal band of hypomera from posteromedial portion to middle of lateral margin, with dispersed but obvious punctures. Elytra. Lateral margins curved outward and elytral surface slightly irregular (rugose). Elytral striae with carinate margins clearly separated. Interstriae flattened or faintly convex with foveiform punctures surrounded by approximately four shiny microtubercles. Microtubercles with sheen more evident than surrounding surface; some microtubercles indistinct. Foveiform punctures separate on the elytral disc by more than one diameter. Basal carina of seventh interstria elongate, length about four times the width of the seventh interestria. Basal carina of ninth interstria with approximately one-half of elytral length. Apical carina or tubercle of second elytral interstria absent. Apical carina of third interstria weaker than apical carina of fourth interstria. Length of apical carina of sixth interstria longer than that of apical carina of fifth interstria. Length of apical carina of seventh interstria shorter than the sum of apical carinae of fifth and sixth interstriae. Pseudepipleural carina complete, visible entire length of epipleuron (as in Fig. 63).

Material studied. Holotype ( BMNH) [examined]: 1. [old white label, printed in black, bordered in red]. Type; 2. [old white label, printed in black]. B. Honduras./ Blancaneau.; 3. [old white label, printed in black]. Sp. figured.; 4. [old white label, handwritten]. Deltochilum acropyge Bates.

Non-type material [2]: GUATEMALA: PETÉN: La Libertad, Comunidad Bethel, 14.VIII.1996, E. Cano (1 ♂ 1 ♀ CEMT).

Sampling methods. baited pitfall trap with live diplopods [2 specimens].

Habitat. Belize, Costa Rica, Guatemala, Mexico and Panama (inhabits the Caribbean slope) [see too Halffter 2003; Kohlmann & Solis, 2012] ( Fig. 91 ), from available data this species inhabits tropical rainforest.

Diagnosis and remarks (within the acropyge complex): Length> 14 mm; posteromedial punctures of pronotum usually separated by at least one diameter; anteromedial punctures of hypomera usually separated by less than one diameter; apex of second interstria without carina or tubercle; pseudepipleural carina complete, visible entire length of epipleuron (as in Fig. 63). Elytra are slightly more convex than in other species of this complex, with clearly impressed striae. The following aedeagus characters are also diagnostic: parameres symmetric and quite elongate, in lateral view forming an angle of approximately 45° with phallobase; apex rounded ( Fig. 13 ). Lamella copulatrix (LC): subrectangular sclerites. Superior right peripheral sclerite (SRP) as in Fig. 46.

Figures

FIGURES 12 – 19. Aedeagus in Deltochilum (Aganhyboma). 12. D. schefflerorum new species. 13. D. acropyge new status. 14. D. larseni new species. 15. D. acanthus. 16. D. longiceps new status. 17. D. valgum. 18. D. icaroides. 19. D. kolleri new species. Figures scale = 1 mm.

FIGURES 1 – 3. Male genital sclerites in Deltochilum (Aganhyboma). 1. Position and nomenclature of endophallic sclerites (modified from Medina et al. 2013). 2. Nomenclature of the main parts of the genital segment (D. amandaarcanjoae new species). 3. Nomenclature of the main parts of the superior right peripheral sclerite (SRP) (D. paresi new species). Figures scale = 1 mm.

FIGURES 103 – 110. Species of D. (Aganhyboma). 103. D. viridescens new status. 104. D. viridicatum new species. 105. D. viridicatum new species. 106. D. acropyge new status. 107. D. feeri new species. 108. D. schefflerorum new species. 109. D. streblopodum new species. 110. D. longiceps new status. Figures scale = 1 mm.

FIGURES 89 – 92. Known distribution of D. (Aganhyboma). 89. trisignatum complex. 90. cupreicolle complex. 91. acropyge complex. 92. finestriatum complex.