Archiinocellia oligoneura Handlirsch, 1910

Archibald, S. Bruce & Makarkin, Vladimir N., 2021, Early Eocene snakeflies (Raphidioptera) of western North America from the Okanagan Highlands and Green River Formation, Zootaxa 4951 (1), pp. 41-79 : 57-58

Archiinocellia oligoneura Handlirsch, 1910

Fig. 11

Type material. Holotype GSC 7250 , collected by L[awrence]. M. L[ambe], 21.vii.1906, deposited in the collections of the GSC, Ottawa. Portions of overlapped fore- and hind wings covered with Canada balsam.

Type locality and horizon. “Opposite Horsefly mine”, Horsefly , British Columbia, Canada; early Eocene (Ypresian) .

Diagnosis. May be distinguished from A. protomaculata by clearly wider wings (forewing length/width 2.69– 3.08 in A. oligoneura ; 3.63–4.15 in A. protomaculata ).

Redescription. Forewing 7–8 mm long (estimated), 2.6 mm wide. Costal space not preserved. ScP very short, terminating on costal margin strongly distad crossvein 2ra-rp; length from termination of ScP to pterostigma slightly greater than length of pterostigma. Distal part of subcostal space broad, with distal crossvein 2scp-r nearly straight. Pterostigma elongate (4.6 times longer than width measured along to costal margin), lightly pigmented. At least two branches of RA: one within pterostigma, strongly inclined, in middle of pterostigma; other forming distal margin of pterostigma; no branch distad pterostigma discernible. RA space with two crossveins forming two radial cells nearly equal in length: 2ra-rp located about mid-way between pterostigma, termination of ScP; 3ra-rp located in distal part of pterostigma along RA. RP originating at obtuse angle, with three pectinate branches: RP1 deeply forked, with anterior branch simple, posterior forked; other branches (RP2, RP3) simple. One long intraradial crossvein rp1-rp2 between stems of RP, RP1. Two preserved between RP, MA (1r-m, 2r-m) connecting stem of RP, MA. M forked well proximad origin of RP, probably fused with CuA for short distance. MA deeply, dichotomously forked twice. MP strongly zigzagged, anterior trace with two simple branches. Two intramedian crossveins form two doi (distal doi markedly longer): 2im very long, joins posterior branch of MA. Anterior trace CuA strongly zigzagged, fused with MP for long distance to form part of posterior margin of basal doi. Crossvein icu between CuA, CuP long. Basal portion, CuP, anal veins not preserved or not clearly understandable.

Hind wing about 6.5–7 mm long (estimated), 2.25 mm wide. Costal space and ScP not preserved. One crossvein detected in subcostal space, 2scp-r, forming basal margin of pterostigma. Distal part of subcostal space broad, with distal crossvein 2scp-r nearly straight. Pterostigma elongate (4.7 times longer than width measured along to costal margin), lightly pigmented. At least two branches of RA: one within pterostigma, strongly inclined, in middle of pterostigma; other forming distal margin of pterostigma; no branches distad pterostigma discernible. RA space with two preserved crossveins: 2ra-rp distad termination of ScP; 3ra-rp located in distal part of pterostigma along RA. RP with three pectinate branches: RP1 deeply forked, with anterior branch simple, posterior forked; other branches (RP2, RP3) simple. One long intraradial crossvein rp1-rp2 between stems of RP, RP1. Two preserved between RP, MA: 2r-m connects stem of RP, MA; 3r-m connects stem of RP1, MA. M probably forked proximad origin of RP. MA deeply dichotomously forked twice. MP: anterior trace zigzagged, simple, with three simple, pectinate branches (alternatively: anterior trace forked, with two simple, pectinate branches). Two intramedian crossveins form two doi, proximal much shorter than distal. One preserved crossvein between M, Cu (2m-cu) connecting MP, CuA. Anterior trace of CuA simple distally, with one preserved simple branch. Basal portion, CuP, anal veins not preserved or not clearly understandable.

Remarks. Handlirsch (1910) incorrectly believed that his Fig. 4 depicted the forewing and his Fig. 5 the hind wing, however, the opposite is true.

Figures

FIGURE 4. Wing venation of Megaraphidia ootsa sp. nov., holotype RBCM P1556. A, right forewing, B, left forewing; C, right hind wing; D, left hind wing. Scale bar = 3 mm (all to scale).

FIGURE 5. Megaraphidia klondika sp. nov., holotype. A, counterpart UWBM 97073; B, part SR03-01-01; C, forewing venation; D, hind wing venation. Scale bar = 3 mm (all to scale).

FIGURE 11. Archiinocellia oligoneura Handlirsch, 1910. A, holotype GSC 7250; B, forewing venation; C, hind wing venation. Venation drawn in B and C overlap on the right half of the photograph, venation on the left side could be of either wing. Scale bars = 3 mm.