Cryptops (Trigonocryptops) Verhoeff, 1906

Schileyko, Arkady A., Vahtera, Varpu & Edgecombe, Gregory D., 2020, An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses, Zootaxa 4825 (1), pp. 1-64 : 20

(!) Cryptops (Trigonocryptops) Verhoeff, 1906

Figs 26, 28, 32

Synonyms. Paratrigonocryptops Demange, 1963

Type species. Cryptops gigas Kraepelin, 1903 (by subsequent designation of Attems, 1930).

Diagnosis. Clypeus with setose plate(s) delimited by sutures ( Fig. 26 , fig. 21 in Schileyko et al. 2018). Pretarsus of maxillae 2 apically pointed, with a ventral projection (as in C. (T.) hephaestus Ázara & Ferreira, 2013 see their fig. 6C) or without it (as in adult (Rc 7502) of C. (T.) sarasini furcatus ( Ribaut, 1923)). Dorsal brush of maxilla 2 as long as or slightly longer than the corresponding pretarsus. Anterior margin of forcipular coxosternite bilobed, in most species with long enlarged marginal setae ( Fig. 26 ); tarsungula long, overlapping each other by at least 1/3 of their length when adducted. Sternites ( Fig. 32 ) with well-developed transverse thickening between coxae of legs; at least some anterior sternites with endosternites bordered by trigonal sutures, the configurations of these sutures may vary (see Schileyko et al. 2018). Anterior corners of the endosternite of some anterior sternites with lateral projections ( Fig. 32 ).

Number of species. 29 ( Schileyko et al. 2018).

Remarks. Treated as a subgenus in Edgecombe & Bonato (2011: 393), Murienne et al. (2011: 62), Ázara & Ferreira (2013: 432), Voigtländer & Reip (2013: 220), Bonato et al. 2016, Lewis (2016a: 575), Schileyko & Stoev (2016: 266), Schileyko et al. (2018: 567).

Cryptops (Paratrigonocryptops) was synonymised to C. (Trigonocryptops) by Lewis (2005). Schileyko & Stoev (2016) considered the similarity between Cryptops (Trigonocryptops) and the nominate subgenus; they reconsidered the recent concept of the former, suggesting that (p. 267) “only species having anterior sternites with complete trigonal sutures and clypeus with setose plate(s) should be assigned to Trigonocryptops ”. Thus “ Cryptops (Trigonocryptops) iporangensis ” Ázara & Ferreira, 2013 should be Cryptops (C.) iporangensis as it has no sternal trigonal sutures. As for Cryptops (Trigonocryptops) similis described by Machado (1953) for four specimens from Southern Spain, it has sternal trigonal sutures that are typical for this subgenus, but there is no information on presence of clypeal setose plate(s) (see pp 85–87) and the corresponding figure III(2) is inadequate. Voigtländer & Reip (2013: 220) mentioned another specimen of Cryptops (T.) similis from Southern Spain but gave no information on the two diagnostic characters of this subgenus (see above). Thus inclusion of Cryptops (T.) similis in Trigonocryptops should be questionable at the moment.

Figures

FIGURES 26–32. Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. paratype CIRAD P8A 26 head, forcipular segment + LBS 1–3 ventrally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 27 forcipular segment ventrally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. holotype MNHN 459 28 right ultimate leg medially; Cryptops (C.) caucasius Verhoeff, 1934; Ad. Rc 8003 29 cephalic plate + anterior margin of forcipular coxosternite ventrally; Cryptops australis (Newport, 1845); AM KS 30 apical part of maxilla 2 laterally; Cryptops (Paracryptops) weberi Pocock, 1891; MCZ DNA 102459 31 apical part of of maxilla 2 laterally; Cryptops (Trigonocryptops) martinicensis Schileyko, Iorio & Coulis, 2018; Ad. holotype MNHN 459 32 LBS 3 ventrally; (am)—anterior margin of forcipular coxosternite, (cl)—clypeus, (cs)—clypeal setose plate, (cst)—forcipular coxosternite, (db)—apical setae of dorsal brush, (es)—enlarged (spiniform) setae, (est)—endosternite, (l)—labrum, (lp)—lateral projection of endosternite, (ls)—long setae, (lsc)—sternal median longitudinal sulcus, (pa)—short rounded projection of anterior margin of forcipular coxosternite, (pf)—prefemur, (sl)—sternal median longitudinal suture, (sr)—sternal transverse ridge, (st)—saw teeth, (sts)—sternal transverse suture, (tgs)—sternal trigonal sutures, (ti)—tibia, (tr)—tarsungula, (vp)—ventral projection of pretarsus of maxilla 2.