Campeprosopa Macquart, 1850

Mason, Franco & Nÿ, Rudolf Rozko Š, 2008, A review of the Oriental Campeprosopa species (Diptera: Stratiomyidae), Zootaxa 1794, pp. 49-64 : 50-51

Campeprosopa Macquart, 1850

Campeprosopa Macquart, 1850: 350 .

Type species: Campeprosopa flavipes Macquart (by original designation)

Diagnosis. Campeprosopa belongs to the subfamily Clitellariinae which is characterised only by a complex of plesiomorphic characters (eight mostly cylindrical antennal flagellomeres, presence of vein M3, crossvein mcu lacking, usually two strong scutellar spines). From other genera of Clitellariinae , Campeprosopa may be distinguished by the antennae inserted on a conical tubercle, by absence of a strong spine in front of the wing base on each side and an elongate abdomen.

Description. The eyes are almost rounded, bare and separated by a frontal vitta in both sexes but touching in the males of C. longispina , facets are small and uniform. The ocellar tubercle is prominent in lateral view, the postocular rim is narrow but usually visible in dorsal view, the occiput is concave. The antennae are placed at about the middle of the head in profile, inserted on a distinct conical protuberance. The basal antennal segments are subequal, or the scape is twice as long as the pedicel, the antennal flagellum is slender and long, consisting of 8 cylindrical but slightly laterally compressed flagellomeres. The lower postocular area is only slightly visible in lateral view. The two-segmented palpus is as long or slightly longer than the half length of the labellum.

The thorax is dark, the thoracic spine above the wing basis is not developed and the scutellum bears a pair of long, slender and diverging spines. The thoracic pile is short, mostly appressed, only the anepisternum partly bare and shining black. The wings may be infuscated to varying extents, but this infuscation is due by the dense microtrichia, which are largely lacking in the central area and the basal third of wing. The anterior crossvein is relatively long and markedly oblique, always shorter than the distance between the crossvein and the origin of R2+3. Vein R4 usually ending in the middle between R2+3 and R5 or somewhat distad. The colour of the legs may be yellow with brown to black patterned (especially in C. flavipes ).

The abdomen is elongated and flattened, parallel-sided or slightly oval and then with maximum width in or beyond the middle, usually dark but pale margined and with more or less extended pale pattern on tergites 2 and 3. Sternites 2 and 3 usually pale.

The male eyes are usually more narrowly separated (or even touching) than in the females, the rim at the inner eye angle is barely distinct or at most twice as broad as lateral ocellus, the frons is relatively narrow (at most about 1.5 times broader than the ocellar tubercle) or reduced. The male antenna is relatively short, only 1.7–3.0 times as long as both basal segments combined. The scape may be as long as or twice as long as the pedicel and the last flagellomere is shorter or longer than the preceding flagellomere. The postpronotal callus on the thorax is more rounded than in the females and the scutellar spines are relatively shorter, not reaching the posterior margin of abdominal segment 2 but conspicuously longer in C. longispina . The abdomen is usually somewhat more slender than in the females, and is parallel-sided. The epandrium is usually as long as proctiger, the cerci are suboval, only slightly shorter than proctiger. The genitalic capsule has a a conical medial process and long gonocoxal apodemes, the gonostylus shows a distict inner lobe apically and long setae on the inner side. The long distal part of the aedeagal complex is bipartite and somewhat dilated apically.

The female frons is usually broader than in the male (2.0–3.0 times than the ocellar tubercle). The flagellum of the female antenna is usually about 3.0–3.3 times as long as both basal antennal segments and the scape and the pedicel are subequal or the scape is twice as long as the pedicel. The last flagellomere is mostly as long as two preceding flagellomeres. The lateral tip of the postpronotal callus is markedly pointed and the scutellar spines are long, almost reaching the posterior margin of abdominal segment 3. The abdomen is usually slightly broader than in the relevant males. In the middle of the abdominal sternite 2 a short cylindrical subconical protuberance is well visible in the females of C. longispina . The female cercus is slender and long, its basal segment being longer than the last abdominal segment and the apical segment only slightly shorter than the basal one.

From our thorough study of material from all the main parts of the Oriental region we concluded that only three species are actually valid and four are synonyms of C. flavipes . According to the present knowledge, the distribution of C. borneensis is confined to the Malaysian part of Borneo, C. flavipes to southermost Thailand, Sumatra and Java (though continental Malaysia is not excluded) and C. longispina is known to occur in India and northern Thailand and Laos.