Triplocania humboldtiana, González-Obando & Carrejo-Gironza & García, 2017

González-Obando, Ranulfo, Carrejo-Gironza, Nancy & García, Alfonso N., 2017, New species of Colombian Triplocania Roesler (Psocodea: ‘ Psocoptera’: Ptiloneuridae), Zootaxa 4336 (1), pp. 1-113 : 55-57

Triplocania humboldtiana n. sp. Male

( Figs 176–181 )

Diagnosis. Belonging in the species assemblage that includes T. calima n. sp., T. felidiaensis n. sp., and T. mariacarmenae n. sp. Differing from them in the hypandrium ( Fig. 180 ) and phallosome ( Fig. 181 ). It has three pairs of endophallic sclerites, the posterior asymmetric, each arm anteriorly curved, right one distally curved, acuminate, left one also distally curved, but apically dilated, with a row of teeth on outer border; mesal pair Ushaped, with arms slender, and a stout median projection bearing distally a row of teeth ( Fig. 181 ).

Color (in 80% ethanol). Body pale cream, with pale brown and dark spots. Compound eyes brown; ocelli hyaline, with ochre centripetal crescents. Head ( Fig. 178 ): with narrow brown diagonal band from inner margin of each compound eye to near the epistomal sulcus. Vertex, front, clypeus and labrum dark cream; postclypeus and vertex with brown spots. Genae with narrow areas brown, from compound eyes to subgenal sulcus. Postgenae cream. Antennae pale brown. Tergal lobes of meso- and metathorax brown. Thoracic pleura cream, except the brown mesopleura. Legs: fore- and hind- pale cream; middle leg pale brown; coxae and tarsomeres 2–3 brown. Forewings with pale brown marginal band from pterostigma to 1A; setae on veins issuing from brown areolae ( Fig. 176 ). Hindwing with cream marginal band, from R4+5 to near Cu1, and anteriorly, along wing margin, from 1A almost to Cu1 ( Fig. 177 ). Abdomen pale cream, with small irregular brown spots. Cluniun brown; hypandrium and phallosome dark brown. Epiproct and paraprocts pale brown.

Morphology. As in diagnosis, plus the following: Head ( Fig. 178 ): H/MxW: 1.49; compound eyes: H/D: 2.32, IO/MxW: 0.75, MxW/IO: 2.27. Vertex almost straight, at the same level of the upper border of the compound eyes. Outer cusp of lacinial tips broad, with seven denticles. Forewings ( Fig. 176 ): pterostigma membrane with abundant setae; L/W: 2.46, pterostigma: lp/wp: 5.50; areola postica high, la/ha: 1.76. Hindwings ( Fig. 177 ): l/w: 2.69. Hypandrium asymmetric ( Fig. 180 ), side sclerites larger than the central one, this deeply convex anteriorly, posterior projection slender, distally acuminate, near the left postero-lateral corner. Phallosome V-shaped anteriorly ( Fig. 181 ), side struts joined distally to the slender external parameres. Paraprocts ( Fig. 179 ) broad, elongate, rounded posteriorly; sensory fields with 37 trichobothria on basal rosettes, setae as illustrated. Epiproct ( Fig. 179 ) wide, semioval, straight anteriorly, rounded posteriorly, with setae as illustrated.

Measurements. FW: 7375, HW: 4975, F: 1725, T: 2925, t1: 1062, t2: 187, t3: 227, ctt1: 26, f1: 1375, IO: 670, D: 570, d: 410, IO/d: 1.63, PO: 0.72.

Material studied. Holotype male. COLOMBIA. Norte de Santander, National Natural Park Tamá, Toledo , La Candelaria ( Pozo Negro ), 7°21’N: 72°28’W, 2203 m., 21–29.XI.2003, C. Leal. Malaise trap ( MAH) .

Etymology. The specific epithet is an adjective; the species is dedicated to the Alexander von Humboldt Biological Resources Research Institute, Boyacá, Colombia.

Figures

FIGURES 176 – 181. Triplocania humboldtiana n. sp. Male. 176. Forewing. 177. Hindwing. 178. Front view of head. 179. Paraprocts and epiproct. 180. Hypandrium. 181. Phallosome. Scales in mm.