Anomala tibialis Lansberge, 1886

Limbourg, Pol, 2016, Contribution to the knowledge of Afrotropical Rutelinae IV (Coleoptera: Scarabaeoidea: Melolonthidae), Belgian Journal of Entomology 38, pp. 1-42 : 1-42

Anomala tibialis Lansberge, 1886

Anomala tibialis LANSBERGE, 1886: 101-102 [described].

Anomala denuda ARROW, 1899: 258-259 [described] New synonymy.

Anomala tibialis OHAUS, 1918: 93 [catalogued]; MACHATSCHKE, 1957: 84 [catalogued]; MACHATSCHKE, 1972: 159 [catalogued]; KRAJCIK, 2007: 50 [catalogued].

Anomala denuda OHAUS, 1918: 90 [catalogued]; BENDERITTER, 1928: 412 [listed]; MACHATSCHKE, 1957: 82 [catalogued]; MACHATSCHKE, 1972: 157 [catalogued]; KRAJCIK, 2007: 31 [catalogued].

TYPE MATERIAL EXAMINED. Holotype ♀ of Anomala tibialis Lansberge, 1886 : [Africa, Congo, Ueth, Evd. Kellen] [Type] [Type] [ Anomala tibialis Lansbe. ] ( RMNH).

Lectotype ♂ of Anomala denuda Arrow, 1899 present designation: [Old Calab., 99 171] [♂] [Syntype] [Type] [ Anomala denuda Arrow Type ♂] [Lectotype ♂ Anomala denuda ARROW, 1899 des. Pol Limbourg, 2004] ( BMNH).

Paralectotype ♀ Anomala denuda Arrow, 1899 : [Old Calab., 99 171] [♀][Syntype] [Type] [ Anomala denuda Arrow Type ♀] [Paralectotype ♀ Anomala denuda Arrow, 1899 det. Pol Limbourg, 2004] ( BMNH).

ADDITIONAL MATERIAL EXAMINED. Senegal: 1♂: Parc du Sine Saloum, Rés. de Fathala , 12.VII.2010, UV, Leg. Alain Coache ( CEREB) ; 1♀: idem, 14.VII.2010 ( CEREB) ; 1♂: idem, 15.VII.2010 ( CEREB) ; 1♀: oriental, Fongoli , 08.IV.2010, UV, Leg. Alain Coache ( CEREB) ; 31♂, 41♀: Ziguinchor, Cap Skirring , 25-28.VII.2008, Leg. P. Moretto ( RBINS) ; Burkina Faso: 1♂: Nasso , X.1995, Leg. D. Camiade ( PCDC) ; 1♀: idem ( RBINS) ; 1♀: Gaoua , 04.VI.1981 ( RBINS) ; Mali: 1♀: Bamako , 01.VII.1988, Leg. J. Hecq ( PCJH) ; 1♂, 1♀: Africa centr., Oubangui-Chari, Leg. Skulina ( NMPC) ; 1♀: Lac Tsad, Leg. Skulina ( NMPC) ; Sierra Leone: 1♀: 13.V.1961, H.M.S. Owen, B.M. 1961-456 ( BMNH) ; Nigeria: 1♀: Centre, nr. Suleja-Abuja (9°11’05N / 7°10’00E), 450m, 19.VII.2006, Leg. P. Léonard & E. Vingerhoedt ( RBINS) ; 1♂, 1♀: Dosso-Moradi, Leg. Skulina ( NMPC) ; 1♀: Sokoto, VII.1976, Leg. C. Joly ( RBINS) ; 1♀: Ogove R., W. Afr., (Good), Holland Coll. ( CMNH) ; Equatorial Guinea: 1♂: Benito W. Afr., Holland Coll. ( CMNH) ; Ivory Coast: 1♀: Mandaloa, Leg. Skulina ( NMPC) ; 1♂, 1♀: Adzopé (06°07’N / 03°51’W), 18.X.2001, Leg. G. Curletti ( MCCI) ; 1♂, 3♀: Adzopé, 04.2002, Leg. Tropini ( MCCI) ; 128specimens: Saiwa , 1995, Leg. D. Camiade ( PCDC) ; 1♂, 1♀: Tonkoui , III.1998, Leg. D. Camiade ( RBINS) ; 10♀: Danane , III.1998, Leg. D. Camiade ( PCDC) ; 8♀: Banco , V.1984, Leg. P. Moretto ( RBINS) ; 1♂: Comoe, IV.1998, Leg. P. Moretto ( RBINS) ; 1♂: Comoe, X.1998, Leg. P. Moretto ( RBINS) ; 1♀: Abidjan, 22.III.1976, light, Leg. J. Hecq ( PCJH) ; 1♀: Abidjan, VI.1980, Leg. P. Moretto ( RBINS) ; 1♀: Abidjan, 28.XII.2010, P. Boireau leg. ( PCPB) ; 1♀: Man , V.2001, Leg. P. Moretto ( RBINS) ; 1♂: Lamto , 07.IV.1981, light ( PCSR) ; 1♀: Lamto , 28.III.1986, light, J. M. Leroux leg. ( PCSR) ; 6♀: Lamto N. R. (6.22°N / 5.03°W), 20- 23.IX.2013, light, T. Delsinne & W. Dekonnick leg. ( RBINS) ; 2♀: Yakassé Mé , P. Boireau leg. ( PCPB) ; Liberia: 1♀: Harbel ( Marshall Terr. ), 05.III.1965, R. M. Fox, Carn. Mus. Acc. 7659 ( CMNH) ; Ghana: 1♂: Central R., Sagiymasse Kibi , X.2003, Leg. P. Legall ( PCLG) ; 2♂, 3♀: Kade, 28.II.1971, light, Collector of John Glaser ( CMNH) ; 3♂, 4♀: Kukurantumi , 13.II.1975, Leg. K. Adlbauer ( PCKW) ; 1♀: Agogo , 09.II.1975, Leg. K. Adlbauer ( PCKW) ; 1♀: Foret de Boabeng , près de Techiman, monkeys sanctuary, 412m, 13.II.2010, UV, Leg. Alain Coache ( CEREB) ; 2♀: Kibi, Fôret de Kibi , 14.II.2010, UV, Leg. Alain Coache ( CEREB) ; Togo: 1♀: Atakpamé , 18.X.1990, Leg. K. Adlbauer ( PCKW) ; 1♀: idem, 21.X.1990 ( PCKW) ; 2♂, 1♀: Rég. des Plateaux, Kloto , VIII.2008, réc. Local collector ( PCSR) ; 1♂, 1♀: Kpalime , 27-30.II.2006, Leg. N. Schiltz ( RBINS) ; Benin: 1♀: Engaradebou , 10.V.2008, Leg. C. Vanderbergh ( CVDB) ; 1♂: Mono, Lokossa, quartier Vivotin , (6°37’03’’N / 1°43’09’’E, 31m), à la lumière, 16.III.2006, Leg. Bernard Rainon ( CEREB) ; 1♀: Mono, Lokossa, quartier Agonvé , (6°38’11’’N / 1°42’28’’E, 40m), à vue, 30.VI.2008, Leg. Bernard Rainon ( CEREB) ; 2♀: Mono, Lobogo, Tokay (6°35'22''N / 1°56'30''E, 30m), ultra-violet, 14.VI.2013, leg. Alain Coache ( CEREB) ; 2♂, 5♀: Mono, Lokossa, Houéyogbé, forêt de Houéyogbé , (6°33'43''N / 1°51'02''E, 82m), ultra-violet, 22.X.2012, Leg. Alain Coache ( CEREB) ; 3♀: idem, 23.IV.2013 ( CEREB) ; 4♂, 3♀: idem, 08.V.2013 ( CEREB) ; 1♂, 1♀: Donga, Bassila, Saramanga, forêt de Goutansoukpa , (9°12’35’’ N / 1°50’16’’E, 341m), à la lumière, 30.V.2012, Leg. Bernard Rainon ( CEREB) ; 1♀: Zou, Zogbodomey, Massi, Forêt de Ko (la Lama ), 24km sud Abomey (2°09’30’’O/ 7°00’04’’N, 51m), 24.I.2010, ultra-violet, Leg. A. Coache ( CEREB) ; 1♀: Zou, Bohicon, quartier Soglogon (7°10'54''N / 2°01'54''E, 200m), ultra-violet, 11.V.2013, Leg. Alain Coache ( CEREB) ; 1♀: Zou, Bohicon, quartier Soglogon , (7°10’54’’N / 2°01’54’’E, 200m), à la lumière, 20.IV.2012, Leg. Bernard Rainon ( CEREB) ; 2♂, 1♀: idem, 24.V.2012 ( CEREB) ; 1♂, 4♀: idem, 01.V.2013, Leg. Alain Coache ( CEREB) ; 4♂: Bohicon, quartier Soglogon , 16.V.2013, ultra-violet, Leg. Bernard Rainon ( CEREB) ; 4♂, 4♀: Donga, Bassila, Saramanga, forêt de Goutansoukpa , (9°12’35’’N / 1°50’16’’E, 341m), à la lumière, 29.IV.2013, Leg. Bernard Rainon ( CEREB) ; 1♀: Donga, Bassila, Pénéssoulou, forêt de Pénélan (9°14'50"N / 1°31'45''E, 392m), 27.V.2013, ultra-violet, Leg. Alain Coache ( CEREB) ; 1♀: idem, 25.V.2013 ( CEREB) ; 1♀: Donga, Bassila, Pénéssoulou, forêt de Pénélan (9°14'50"N / 1°31'45''E, 392m), 22.V.2013, ultra-violet, Leg. Alain Coache ( CEREB) ; 1♂, 6♀: Donga, Bassila, Pénéssoulou , stade foot (9°14’15’’N / 1°33’12’’E, 415m), 15.V.2013, ultra-violet, Leg. Alain Coache ( CEREB) ; 5♂, 5♀: B. Atlantique, Ouidah, forêt de Pahou ( Ahozon ) (6°23'03''N / 2°09'32''E, 29m), 09.V.2013, Leg. Bernard Rainon ( CEREB) ; 1♀: Mono, Athiémé , bord du Mono (6°14'20''N / 1°40'00''E, 29m), ultra-violet, 31.V.2013, Leg. Alain Coache ( CEREB) ; 3♀: Donga, Bassila, Saramanga village (9°13'09''N / 1°46'29''E, 392m), 24.V.2013, ultra-violet, Leg. Alain Coache ( CEREB) ; 2♀: Ouidah , foret de Pahou, 09.V.2013, ultra violet, Leg. Bernard Rainon ( CEREB) ; 2♂: Houeyogbé , foret de Houeyogbé, 08.V.2013, ultra violet, Leg. Bernard Rainon ( CEREB) ; 1♂: Goutansoukpa , light trap, forest area, VI.2007, Col. G. Goergen ( IITA) ; Cameroon: 1♂, 1♀: Bangangte W., 26.II.1993, Leg. M. Baffray ( PCAT) ; 1♀: Obout , leg. J. M. M’Bida ( PCAT) ; 1♂: Alantika Mts., Wangou , V.2010, Leg. P. Legall ( PCLG) ; 1♂: South, Mt. Koupe, Nyassoso , 900m, XII.2007, Leg. P. Legall ( PCLG) ; 1♀: Sangmelima , X.1933, A. I. Good, Carn. Mus. Acc. 19080 ( CMNH) ; 1♂: Lolodorf , 15.III.1923, A. I. God, Carn. Mus. Acc. 7295, 526 ( CMNH) ; 2♂: Ndom, N. Sanaga , II.1995, Leg. P. Moretto ( RBINS) ; 1♂, 1♀: Nyassosso , I-III.2008, Leg. P. Legall ( PCLG) ; 2♀: Duala ( NMPC) ; Gabon: 1♀: Ntoum , IX.1994, Leg. A. Pauly ( RBINS) ; 2♂, 1♀: Ogove R., W. Afr. (Good), Holland collection ( CMNH) ; 1♂: Libreville, Arboretum , XI.2009, Leg. Ph. Oremans ( PCPO) ; 4♂, 10♀: La Poppée, IX.2009, Leg. Ph. Oremans ( PCPO) ; Republic of Congo: 4♀: Brazzaville, Leg. Vosahle ( NMPC) ; 7♀: Mayombé, Tessi , XII.1992, Leg. P. legall ( PCLG) ; 4♀: Tsessi ( Mayombe ), XI.1992, Leg. D. Camiade ( PCDC) ; 1♀: idem, XI.1993 ( PCDC) ; 2♂: Mayombé, Dimonika , XII.1990, Leg. P.Legall ( PCLG) ; 1♂, 1♀: Voka (S 4°38’E 14°40‘), X-XI.2009, Leg. E. Vingerhoedt ( RBINS) ; 1♂, 3♀: N’Goma , P.N. du Conkouati, 17.V.2010, UV, Leg. Alain Coache ( CEREB) ; 6♀: N’Goma , P.N. du Conkouati, 25.V.2010, sur fleurs Albizia, Leg. Alain Coache ( CEREB) ; 3♀: N’Goma , P.N. du Conkouati, 02.VI.2010, UV, Leg. Alain Coache ( CEREB) ; 1♀: Kouilou, Conkovati , 01.III.1997, Leg. D. Camiade ( RBINS) ; 4♀: Mayombé, Dimonika , XI.1991, Leg. P. Legall ( PCLG) ; 6♂, 3♀: Diba ( Mbomo ), II.1991, Leg. D. Camiade ( PCDC) ; 3♀: Diba , II.1991, Leg. P. Legall ( PCLG) ; 2♂, 1♀: Kinkengue , XI.1992, Leg. D. Camiade ( PCDC) ; 1♂, 3♀: Omoy ( Chaillu Mts. ), XI.1992, Leg. D. Camiade ( PCDC) ; 1♂: Etoumbi , XII.1993, Leg. D. Camiade ( PCDC) ; 1♂: idem, II.1994 ( PCDC) ; 1♀: Mabaya , X.1990, Leg. P. Legall ( PCLG) ; 1♂: Mah, Plateau Tekes , XI.1991, Leg. P. Legall ( PCLG) ; 1♂: Linzolo , 09.XI.1992, Leg. C. Vanderbergh ( CVDB) ; 1♂, 1♀: Dimonika , XI.1992, Leg. C. Vanderbergh ( CVDB) ; ♀: Ikefemba , Carn. Mus. Acc. 8852 ( CMNH) ; Democratic Republic of Congo: 1♂, 1♀: ABIO (S3°07’/E15°31’), Lesio-Louna Réserve , XI.2009, Leg. E. Vingerhoedt ( RBINS) ; 5♂, 5♀: Kiwu-Ituri, Leg. Skulina ( NMPC) ; 1♀: Beni ( Kivu ), IV.1977, Coll. R. Ducarme ( PCRD) ; 1♂, 2♀: idem, 1980 ( PCRD) ; 1♀: idem, 1985 ( PCRD) ; 1♂: idem, 1986 ( PCRD) ; 1♂: Kinshasa, Tilapia , 09.VI.1979, Leg. C. Joly ( RBINS) ; 2♀: idem, X.1979 ( RBINS) ; 2♀: idem, 09.XI.1983 ( RBINS) ; 1♀: idem, 10.XI.1983 ( RBINS) ; 2♂: Bombo Lumene, rég. Kinshasa , III.1996, Leg. P. Oremans ( RBINS) ; 1♂: Likouala region, Epena (02°34’S / 17°24’E), 340m, 11-13.VI.1993, J. Rawlins, G. Onore, R. Davidson & D. Schlitter ( CMNH) ; 1♂: Likouala region, Impfondo (02°26’S / 18°00E), 340m, 09- 11.VI.1993, J. Rawlins, G. Onore, R. Davidson & D. Schlitter ( CMNH) ; 1♂: Benito W. Afr., Holland Collection ( CMNH) ; 11♀: Nord Kivu , env. Beni, IV-V.2009, Leg. R. Ducarme ( RBINS) ; 1♀: Equateur Prov., Lukolela zone, Bobeta, X-XII.2007, Leg. Ph. Oremans ( PCPO) ; 1♀: Mbuji-Mayi , XII.1979, Leg. C. Joly ( RBINS) ; Zambia: 1♀: Changala, Leg. Skulina ( NMPC) .

COMMENTS

After examination of types of A. tibialis Lansberge, 1886 and A. denuda Arrow, 1899 it appears those are conspecific.

DISTRIBUTION. Widespread, currently known from Senegal to Democratic Republic of Congo and Zambia.

BIOLOGY. Specimens from Republic of Benin were collected in secondary forest (pers. com. A. Coache).

The species was collected from January to December.