Tetramorium chefketi
publication ID |
https://doi.org/ 10.5281/zenodo.399638 |
DOI |
https://doi.org/10.5281/zenodo.5629590 |
persistent identifier |
https://treatment.plazi.org/id/03DB878C-3704-7E47-FF70-6FA9FF40EE93 |
treatment provided by |
Plazi |
scientific name |
Tetramorium chefketi |
status |
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Definition of chefketi species complex
Species belonging to the T. chefketi species complex can be defined by the following combination of characters:
Workers
a) polygonal striae are continuous on 1st gastral tergite, and is sometimes microreticulate basally. Polygonal striation: interstices marked by feeble anastomosing striae (Fig. 8.), compare to microreticulation: interstices marked by very feeble elevated structures, like rugulae (Fig. 6, 7.);
b) head, alitrunk and waist coarsely sculptured, dorsum of petiolar node and postpetiole coarsely rugose and/ or reticulate.
c) petiole and postpetiole relatively narrow (CS/PEW and CS/PPW, Table 1 View TABLE 1 );
d) ground surface on the frons is usually coarsely microreticulate;
Gynes
e) alitrunk is low, dorsum is flattened in profile;
f) scutum in dorsal view is somewhat narrowed anteriorly and does not cover completely the pronotum, humeri are well visible;
g) head, alitrunk and waist are coarsely sculptured, and petiole and postpetiole dorsum coarsely rugose or/ and reticulate;
h) polygonal striae disrupted on 1st first gastral tergite, sometimes microreticulate basally;
i) mandibles are longitudinally striate;
j) petiole and postpetiole are relatively narrow (WAIST, Table 2 View TABLE 2 );
Males
k) head and alitrunk with usually coarsely sculpture, ground surface microreticulate; l) sides of petiole and postpetiole not angulate as in the ferox species complex (in dorsal view); m) first gastral tergite usually smooth and shiny, or feebly striate;
n) stipes of genitalia blunt, curved inwards at the top, with flattened apical plate (in caudal view);
We placed the following species into the chefketi species complex:
1) Tetramorium alternans Santschi, 1929
Tetramorium biskrensis kahenae Menozzi, 1934 new synonymy
2) Tetramorium anatolicum Csősz & Schulz , new species 3) Tetramorium annectens Pisarski, 1969 status revised
alternans anatolicum annectens chefketi moravicum sanetrai sulcinode (n=5) (n=3) (n=1) (n=11) (n=12) (n=3) (n=3) 4) Tetramorium chefketi Forel, 1911
Tetramorium caespitum View in CoL var. sarkissiani Forel, 1911 new synonymy Tetramorium turcomanicum Santschi, 1921 View in CoL new synonymy Tetramorium taurocaucasicum Arnoldi, 1968 new synonymy Tetramorium forte Forel, 1904 View in CoL [in part, only material from Crimea and Caucasus] (for details of
clarification of synonymy see Taxonomic notes under Tetramorium chefketi )
5) Tetramorium exile Csősz & Radchenko , new species
6) Tetramorium moravicum Kratochvíl, 1941
Tetramorium rhenanum Schulz, 1996
7) Tetramorium rhodium Emery, 1922 status revised
8) Tetramorium sanetrai Schulz & Csősz , new species
9) Tetramorium sulcinode Santschi, 1927 status revised
Tetramorium karakalense Dlussky & Zabelin, 1985 View in CoL new synonymy
10) Tetramorium syriacum Emery, 1922 View in CoL
The newly defined chefketi species complex may not strictly reflect the phylogenetic relationship of the abovementioned species. T. sulcinode and T. annectens possess several autapomorphic characters (e.g. relatively large eyes and a long psammophore) what probably may lead to place them into a separate group in a future.
alternans (n=34) | anatolicum (n=38) | annectens chefketi exile moravicum rhodium (n=8) (n=131) (n=33) (n=108) (n=22) | sanetrai sulcinode (n=15) (n=17) | syriacum View in CoL (n=23) | |
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CS | 733 ±35.37 [652, 820] | 702 ±23.87 [635, 743] | 675 ±17.39 869 ±51.36 696 ±23.95 833 ±60.74 832 ±52.91 [645, 703] [740, 972] [620, 742] [720, 953] [720, 895] | 740 ±31.90 801 ±31.65 [683, 783] [773, 853] | 832 ±16.18 [807, 865] |
CL/CW | 1.01 ±0.01 [0.97,1.05] | 1.02 ±0.02 [0.99, 1.05] | 1.02 ±0.02 1.01 ±0.02 1.06 ±0.01 1.02 ±0.02 0.99 ±0.01 [1.0, 1.05] [0.97, 1.04] [1.04, 1.08] [0.97, 1.06] [0.98, 1.02] | 1.01 ±0.02 1.02 ±0.02 [0.98, 1.04] [0.98, 1.03] | 1.02 ±0.01 [0.99, 1.04] |
FR/CS | 0.39 ±0.01 [0.37,0.41] | 0.37 ±0.01 [0.35, 0.39] | 0.39 ±0.01 0.37 ±0.01 0.34 ±0.01 0.36 ±0.01 0.38 ±0.01 [0.38, 0.40] [0.35, 0.39] [0.33, 0.37] [0.34, 0.38] [0.36, 0.40] | 0.37 ±0.01 0.37 ±0.01 [0.36, 0.38] [0.36, 0.38] | 0.50 ±0.01 [0.49, 0.55] |
FL/FR | 1.00 ±0.00 [1.0, 1.0] | 1.02 ±0.05 [1.0, 1.08] | 1.02 ±0.01 1.03 ±0.02 1.0 ±0.01 1.01 ±0.01 1.0 ±0.0 [1.0, 1.04] [1.0, 1.09] [1.0, 1.02] [1.0, 1.04] [1.00, 1.02] | 1.00 ±0.00 1.01 ±0.01 [1.0, 1.02] [1.0, 1.02] | 1.01 ±0.01 [1.00, 1.02] |
SL/CS | 0.71 ±0.01 [0.69,0.74] | 0.78 ±0.01 [0.76, 0.81] | 0.76 ±0.01 0.82 ±0.02 0.84 ±0.01 0.80 ±0.02 0.73 ±0.01 [0.73, 0.77] [0.78, [0.82, 0.85] [0.77, 0.83] [0.71, 0.75] 0.87] | 0.74 ±0.02 0.78 ±0.01 [0.72, 0.75] [0.76, 0.81] | 0.75 ±0.01 [0.70, 0.77] |
EYE* | 0.168±0.01 [0.155, 0.176]* | 0.176 ±0.0 [0.167, 0.182]* | 0.205±0.01 0.171±0.01 0.167±0.00 0.170±0.01 0.173±0.01 [0.196, [0.165, [0.158, [0.165, [0.163, 0.214]* 0.184]* 0.175]* 0.183]* 0.184]* | 0.172±0.01 0.186±0.01 [0.164, [0.180, 0.185]* 0.190]* | 0.162±0.01 [0.155, 0.171]* |
ML/CS | 1.10 ±0.03 [1.04,1.14] | 1.08 ±0.03 [1.06, 1.16] | 1.13 ±0.04 1.18 ±0.03 1.14 ±0.02 1.18 ±0.03 1.12 ±0.02 [1.06, 1.16] [1.10, 1.23] [1.10 1.21] [1.13, 1.26] [1.08, 1.16] | 1.13 ±0.02 1.12 ±0.02 [1.10, 1.18] [1.09, 1.17] | 1.16 ±0.02 [1.09, 1.20] |
MW/CS | 0.63 ±0.01 [0.60,0.65] | 0.62 ±0.01 [0.60, 0.65] | 0.65 ±0.02 0.64 ±0.02 0.63 ±0.01 0.64 ±0.02 0.64 ±0.01 [0.62, 0.68] [0.60, 0.68] [0.61, 0.65] [0.59, 0.69] [0.62, 0.66] | 0.63 ±0.01 0.65 ±0.01 [0.62, 0.65] [0.63, 0.67] | 0.65 ±0.01 [0.63, 0.66] |
PEW/ PPW | 0.86 ±0.04 [0.80,0.91] | 0.80 ±0.02 [0.74, 0.83] | 0.73 ±0.02 0.85 ±0.03 0.86 ±0.03 0.83 ±0.03 0.84 ±0.02 [0.70, 0.75] [0.76, 0.93] [0.80, 0.92] [0.75, 0.89] [0.71, 0.90] | 0.83 ±0.02 0.84 ±0.01 [0.78, 0.85] [0.82, 0.85] | 0.84 ±0.02 [0.81, 0.88] |
NOH/ NOL | 1.03 ±0.06 [0.94,1.13] | 1.15 ±0.08 [1.0, 1.30] | 1.10 ±0.11 0.86 ±0.06 1.04 ±0.06 1.07 ±0.08 0.84 ±0.03 [0.96, 1.25] [0.76, 0.97] [0.90, 1.17] [0.90, 1.29] [0.76, 0.97] | 0.89 ±0.04 1.14 ±0.06 [0.83, 0.97] [1.06, 1.21] | 0.84 ±0.03 [0.78, 0.89] |
PEH/ NOL | 1.84 ±0.12 [1.68,2.04] | 2.01 ±0.08 [1.74, 2.30] | 2.06 ±0.08 1.52 ±0.10 1.83 ±0.06 1.94 ±0.12 1.50 ±0.04 [1.96, 2.12] [1.37, 1.69] [1.69, 1.96] [1.72, 2.23] [1.38, 1.73] | 1.54 ±0.04 1.99 ±0.07 [1.47, 1.63] [1.87, 2.07] | 1.51 ±0.04 [1.43, 1.6] |
NOL/ PEL | 0.87 ±0.07 [0.73,1.00] | 0.76 ±0.06 [0.67, 0.93] | 0.78 ±0.05 0.98 ±0.05 0.86 ±0.03 0.85 ±0.05 1.01 ±0.03 [0.67, 0.83] [0.81, 1.12] [0.80, 0.94] [0.77, 0.94] [0.94, 1.06] | 1.03 ±0.06 0.83 ±0.03 [0.95, 1.12] [0.80, 0.88] | 1.01 ±0.03 [0.97, 1.08] |
CS/ PEW | 3.11 ±0.13 [2.93,3.39] | 3.49 ±0.14 [3.22, 3.75] | 3.50 ±0.19 3.16 ±0.15 3.32 ±0.14 3.18 ±0.14 3.29 ±0.11 [3.35, 3.81] [2.72, 3.49] [3.09, 3.69] [2.84, 3.50] [3.07, 3.53] | 3.05 ±0.09 3.17 ±0.12 [2.91, 3.20] [3.03, 3.33] | 3.26 ±0.11 [3.05, 3.52] |
CS/ PPW | 2.66 ±0.11 [2.42,2.85] | 2.79 ±0.10 [2.56, 3.03] | 2.55 ±0.13 2.70 ±0.13 2.84 ±0.09 2.63 ±0.09 2.80 ±0.06 [2.38, 2.80] [2.33, 2.92] [2.64, 3.01] [2.40, 2.88] [2.64, 3.0] | 2.52 ±0.08 2.67 ±0.09 [2.38, 2.74] [2.58, 2.82] | 2.75 ±0.06 [2.65, 2.88] |
CS | 999 ±31.35 [948, 1025] | 1012 ±38.84 [980, 1055] | 997.5 | 1121 ±38.97 [1060, 1180] | 1198 ±79.59 [1045, 1275] | 986 ±11.81 [973, 995] | 973 ±39.9 [940, 1018] |
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CL/CW | 0.92 ±0.02 [0.90, 0.95] | 0.93 ±0.04 [0.90, 0.98] | 0.98 | 0.92 ±0.02 [0.89, 0.95] | 0.88 ±0.02 [0.84, 0.92] | 0.89 ±0.02 [0.86, 0.90] | 0.94 ±0.02 [0.92, 0.97] |
FR/CS | 0.41 ±0.01 [0.40, 0.42] | 0.37 ±0.02 [0.36, 0.40] | 0.39 | 0.38 ±0.01 [0.36, 0.40] | 0.36 ±0.01 [0.34, 0.38] | 0.38 ±0.00 [0.37, 0.38] | 0.37 ±0.01 [0.36, 0.37] |
FL/FR | 1.00 ±0.00 [1.0, 1.0] | 1.01 ±0.03 [1.0, 1.03] | 1.0 | 1.01 ±0.01 [1.0, 1.03] | 1.0 ±0.01 [1.0, 1.02] | 1.00 ±0.00 [1.0, 1.0] | 1.00 ±0.00 [1.0, 1.0] |
SL/CS | 0.69 ±0.02 [0.65, 0.71] | 0.75 ±0.03 [0.71, 0.77] | 0.72 | 0.77 ±0.01 [0.75, 0.79] | 0.73 ±0.01 [0.71, 0.75] | 0.67 ±0.01 [0.66, 0.68] | 0.73 ±0.01 [0.72, 0.74] |
MW/CS | 0.94 ±0.02 [0.92, 0.97] | 0.96 ±0.05 [0.92, 1.01] | 0.92 | 0.94 ±0.02 [0.88, 1.01] | 1.07 ±0.02 [1.05, 1.13] | 0.94 ±0.01 [0.94, 0.95] | 0.93 ±0.01 [0.92, 0.94] |
PEW/PPW | 0.76 ±0.01 [0.74, 0.77] | 0.72 ±0.04 [0.68, 0.75] | — | 0.78 ±0.03 [0.75, 0.82] | 0.71 ±0.03 [0.66, 0.76] | 0.76 ±0.02 [0.73, 0.77] | 0.75 ±0.01 [0.74, 0.76] |
NOH/NOL | 1.44 ±0.14 [1.31, 1.61] | 1.70 ±0.25 [1.41, 1.86] | 1.38 | 1.14 ±0.12 [1.0, 1.31] | 2.07 ±0.21 [1.69, 2.42] | 1.35 ±0.10 [1.28, 1.47] | 1.51 ±0.17 [1.38, 1.70] |
NOH/PEL | 0.87 ±0.10 [0.73, 0.98] | 0.95 ±0.07 [0.87, 1.0] | 0.94 | 0.83 ±0.09 [0.71, 0.93] | 0.90 ±0.07 [0.82, 1.0] | 0.91 ±0.01 [0.90, 0.92] | 0.92 ±0.08 [0.85, 1.0] |
NOL/PEL | 0.60 ±0.04 [0.56, 0.65] | 0.56 ±0.05 [0.53, 0.62] | 0.68 | 0.73 ±0.06 [0.60, 0.82] | 0.44 ±0.05 [0.34, 0.52] | 0.68 ±0.05 [0.63, 0.72] | 0.61 ±0.02 [0.59, 0.62] |
PEH/NOL | 2.89 ±0.13 [2.75, 3.04] | 3.04 ±0.47 [2.53, 3.46] | 2.56 | 2.18 ±0.17 [2.0, 2.54] | 3.95 ±0.36 [3.31, 4.50] | 2.58 ±0.19 [2.44, 2.80] | 2.77 ±0.33 [2.53, 3.15] |
PEW/PEH | 0.94 ±0.05 [0.86, 0.98] | 0.86 ±0.03 [0.83, 0.88] | 0.80 | 0.94 ±0.03 [0.89, 0.99] | 0.89 ±0.04 [0.83, 0.94] | 0.96 ±0.07 [0.89, 1.02] | 0.82 ±0.01 [0.82, 0.83] |
CS/PEW | 2.49 ±0.20 [2.31, 2.83] | 2.82 ±0.15 [2.72, 2.99] | 2.85 | 2.52 ±0.06 [2.43, 2.60] | 2.61 ±0.16 [2.33, 2.80] | 2.37 ±0.06 [2.31, 2.43] | 2.86 ±0.05 [2.81, 2.91] |
CS/PPW | 1.89 ±0.13 [1.76, 2.11] | 2.01 ±0.04 [1.97, 2.04] | — | 1.97 ±0.08 [1.82, 2.11] | 1.86 ±0.10 [1.74, 2.02] | 1.79 ±0.03 [1.77, 1.82] | 2.15 ±0.06 [2.09, 2.21] |
WAIST | 0.93 ±0.06 [0.83, 1.0] | 0.85 ±0.02 [0.83, 0.87] | — | 0.90 ±0.03 [0.86, 0.96] | 0.92 ±0.05 [0.86, 1.0] | 0.98 ±0.01 [0.97, 0.99] | 0.82 ±0.02 [0.80, 0.84] |
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Kingdom |
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Phylum |
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Class |
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Order |
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Family |
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Genus |
Tetramorium chefketi
Sz, Sándor Cs Ő, Radchenko, Alexander & Schulz, Andreas 2007 |
Tetramorium rhenanum
Schulz 1996 |
Tetramorium karakalense
Dlussky & Zabelin 1985 |
Tetramorium taurocaucasicum
Arnoldi 1968 |
Tetramorium biskrensis kahenae
Menozzi 1934 |
Tetramorium rhodium
Emery 1922 |
Tetramorium syriacum
Emery 1922 |
Tetramorium turcomanicum
Santschi 1921 |
Tetramorium chefketi
Forel 1911 |
var. sarkissiani
Forel 1911 |
Tetramorium forte
Forel 1904 |