Reddyanus Vachon, 1972

Kovařík, František, Lowe, Graeme & Šťáhlavský, František, 2020, Reddyanus justi sp n from Laos (Scorpiones Buthidae), Euscorpius 321, pp. 1-11 : 1-3

Reddyanus Vachon, 1972

( Figures 1–51 , Table 1 )

http://zoobank.org/urn:lsid:zoobank.org:act:02B94E0F- 4071-4447-A8E3-04647C934E39

Isometrus (Reddyanus) Vachon, 1972: 176–177 , figs. 14, 16, 18, 20; Vachon, 1982: 90–101, 108–110, figs. 47–67, 88–91; Fet & Lowe, 2000: 150–154; KovařÍk, 2003: 5–14, figs. 1–8; KovařÍk & Ojanguren, 2013: 184–193, 347–354, 357–360, figs. 1265–1367, 1384–1409.

Isometrus (Raddyanus [sic]): Tikader & Bastawade, 1983: 254–311 (in part), figs. 727–770, 841–895.

Reddyanus KovařÍk et al., 2016: 53–88 , figs. 14, 199, 201– 220, 225–236, 245–250, 254–402, 409–420, 551–554, 561–570, tables 3–5; KovařÍk & ŠťÁhlavský, 2019: 1–45, figs. 1–226, tables 1–4.

TYPE SPECIES. Isometrus acanthurus Pocock, 1899

DIAGNOSIS. Medium sized buthids, adults 19–75 mm. Carapace anteriorly emarginate, surface granulose, carinae obsolete eXcept for anterior medians in some species. Three to five pairs of lateral eyes. Cheliceral fiXed finger with a single ventral denticle. Sternum type 1, triangular in shape. Pectines with fulcra. Tergites with one median carina. Telson with subaculear tooth pointed or rounded. Pedipalps orthobothriotaXic, type Aβ. Trichobothrium d 2 of femur located internally, d

3 of patella located between dorsomedian and dorsoeXternal carina, db of chela located between et and est. Pedipalp movable finger with siX rows of granules, several accessory granules and eXternal and internal accessory granules. Pedipalp fiXed finger with seven rows of granules and siX eXternal and seven internal accessory granules. Legs III–IV without tibial spurs. Tibia and tarsomeres of legs I–III with setae not arranged into bristle combs on dorsal surfaces. Tarsomere II of leg IV with two sparse rows of <20 spiniform setae on ventral surface. Males of most species with more elongate metasomal segments, and often also a wider pedipalp chela manus than females.

SUBORDINATE TAXA. Currently includes 33 species (see KovařÍk & ŠťÁhlavský, 2019: 3).

DISTRIBUTION. From India and China (Tibet) in the west to Melanesia in the east. For distribution in Southeast Asia see Fig. 51 .

Figures

Figure 51. Map showing confirmed distribution of Reddyanus spp. in Southeast Asia.

Figures 1–2. Reddyanus justi sp. n., male holotype (1) and male holotype (right) with female paratype (2) in vivo habitus.

Figures 3–9: Reddyanus justi sp. n. Figures 3–4, 8–9. Male holotype in dorsal (3) and ventral (4) views, and telson lateral (whole view, 8 and detail for subaculear tubercle, 9). Figures 5–7. Female paratype in dorsal (5) and ventral (6) views, and telson lateral (7) view. Scale bar: 10 mm (3–6).

Figures 10–15: Reddyanus justi sp. n., metasoma and telson. Figures 10–12. Holotype male, lateral (10), ventral (11), and dorsal (12) views. Figures 13–15. Paratype female, lateral (13), ventral (14), and dorsal (15) views. Scale bars: 10 mm (10–12, 13–15).

Figures 16–23: Reddyanus justi sp. n. Figures 16–17, 20–23. Male holotype, carapace and tergites I–III (16), coXosternal area and sternites (17), and left legs I–IV, retrolateral aspect (20–23). Figures 18–19. Female paratype, carapace and tergites I–III (18) and coXosternal area and sternites (19).

Figures 24–43: Reddyanus justi sp. n. Figures 24–34. Male holotype, pedipalp chela dorsal (24), eXternal (25) and ventral (26) views, patella dorsal (27), eXternal (28) and ventral (29) views, femur and trochanter dorsal (30), internal (31), and ventral (29) views, movable finger (33) and fiXed finger (34) dentition. Figures 35–43.Female paratype, pedipalp chela dorsal (35), eXternal (36) and ventral (37) views, patella dorsal (38), eXternal (39) and ventral (40) views, femur and trochanter dorsal (41), internal (42), and ventral (43) views. Trichobothrial pattern is indicated in Figures 24–28 and 30–31.

Figures 44–48: Reddyanus justi sp. n., left hemispermatophore. Figures 44–45. Capsule and flagellum, conveX (44) and posterior-conveX (45) views. Figures 46–48. Whole hemispermatophore, conveX (46), posterior-conveX (47) and concave (48) views. Scale bars: 500 μm (44–45), 1 mm (46–48).

Figures 49–50. Reddyanus justi sp. n., male holotype. Postpachytene (49) and ideogram (50) (y aXis - % of the diploid chromosome length, dark grey marks chromosomes in multivalent association). Arrowhead indicates a slightly longer pair of chromosomes. Scale bar: 5 μm (49).