Coeligetes howardi, Bezděk, 2016

Bezděk, Jan, 2016, Revision of the genus Coeligetes from Malaysia and Indonesia, and description of Coeligetoides gen. nov. (Coleoptera: Chrysomelidae: Galerucinae), Zootaxa 4085 (4) : -

Coeligetes howardi BezdĚK, SP. nov.

( Figs 26–27 , 35–42 )

Coeligetes unicolor: Mohamedsaid, 1994: 86 ; Mohamedsaid, 1995b: 5; Mohamedsaid, 1997a: 164; Mohamedsaid, 1997b: 202; Mohamedsaid, 1998b: 258; Mohamedsaid, 1999b: 122; Mohamedsaid, 1999c: 15; Mohamedsaid & Holloway, 1999: 165; Mohamedsaid, 2000: 310; Mohamedsaid, 2004: 101 (catalogue).

Type locality. Indonesia, Central Kalimantan, Busang and Rekut rivers confluence, 0°03′S 113°59′E.

Type material. Holotype: ♂ (BMNH), “INDONESIA: Borneo / Kalimantan Tengah / Busang / Rekut confl. / 0°03 ' S, 113°59 ' E [w, p] // August 2001 / MV light / Brendell/Mendel [w, p] // „Barito Ulu 2001“ / BMNH(E) / 2001-191 [w, p]”. Paratypes: 3 ♀ (BMNH), same data as in holotype; 1 ♂ (JBCB), “BORNEO, SABAH, 15-27.4. / Crocker Mt., 1993 / Gunong Emas env. / I. Jeniš leg. [w, p]”; 1 ♂ 17 ♀ (BMNH), “BORNEO, Sabah / Crocker Range / 26.iii. 2007, 700m / Steven Chew [w, p] // BMNH{E} / 2009-6 [w, p]”; 4 ♀ (BMNH), “BORNEO, Sabah / Mount Trus Madi, / 20.iii. 2007, 600m / Steven Chew [w, p] // BMNH{E} / 2009-6 [w, p]”; 2 ♂ 1 ♀ (PRCS), “MALAYSIA, N Borneo, Sabah, / Keningau dist., Trus Madi Mt., / 1250m, N 05°26 ' 35 '', E 116°27 ' 5 '' / 17– 27.III.2012 P. Romantsov leg. [w, p]”; 3 ♂ 43 ♀ (PRCS), same data, but 16–27.III.2012; 1 ♀ (PRCS), same data, but 17.III.2012; 1 ♀ (PRCS), same data, but 20–22.III.2012; 1 ♂ 1 ♀ (PRCS), same data, but 27.III.2012; 2 ♂ 9 ♀ (PRCS), same data, but 4.IV.2013; 4 ♂ 10 ♀ (PRCS), same data, but 6.IV.2013; 5 ♀ (PRCS), same data, but 9.IV.2013; 1 ♂ (PRCS), “MALAYSIA, S Borneo, Sabah, / Nabawan dist., ~ 7 km / N Pensiangan vill., h~ 530m / N 04°35 ' 16 '', E 116°19 ' 27 '' / 27–31.V.2014 A. Klimenko leg. [w, p]”; 1 ♂ (PRCS), “MALAYSIA,Tambunan, Sabah, / Borneo Is. 20.VII.–20.VIII.2011 / A. Tsylin leg. [w, p]”; 1 ♂ (PRCS), “Borneo Isl, Sabah, Keningau / district, Tambunan h~ 570 m / N 0 5.40861 / E 116.22523 / Pushenkov A. leg. 07-09. [w, p]”; 1 ♀ (BMNH), “BORNEO, Sabah / Sandakan, / 13.ii.2007 / Steven Chew [w, p] // BMNH{E} / 2009-6 [w, p]”; 2 ♀ (BMNH), “BORNEO, Sabah / Ranau, / 13.iii. 2007, 800m / Steven Chew [w, p] // BMNH{E} / 2009-6 [w, p]”; 2 ♀ (BMNH), “BORNEO, Sabah / Ranau, 5.iv. 2007, 500m / Steven Chew [w, p] // BMNH{E} / 2009-6 [w, p]”; 1 ♀ (JBCB), “SARAWAK: Lanjak / Entimau / 28–29 Feb. 1992 Zaidi [w, p] // Coeligetes / unicolor Jac. [h] / det. Mohamedsaid 199 [p] 7 [w, h]”; 1 ♀ (NMPC), “INDONESIA, E Kalimantan / ca. 55 km W of Balikpapan / PT Fajar Surya Swadaya (area) / 01°16.4 ' S, 116°21.1 ' E, 82 m / J. Hájek, J. Schneider & / P. Votruba leg. 23.xi.–1.xii.2011 [w, p] // base camp surrounded with / Acacia plantation; individual / collecting on vegetation, dead / wood and in puddles, + light trap [w, p]”; 1 ♀ (BMNH), “BORNEO, Sabah / Mount Trus Madi / 14-viii.2005 / Steven Chew leg. / BMNH(E) 2006- 36 [w, p]”; 1 ♀ (BMNH), “Primitive / forest. [w, p] // Beating undergrowt. [w, p] // SARAWAK: / Mt. Dulit, / R. Koyan 2,500 ft. / Primary forest. [p] / 18. [h] xi.1932. [w, p] // Oxford Univ. Exp. / B. M. Hobby & / A. W. Moore. / B. M. 1933-254. [w, p]”; 1 ♀ (BMNH), “B.N. BORNEO. / Mt. Kinabalu, / Kiau, 3,000 ft. [p] / 8. 4. [h] 1929. [pink label, p] // Ex F. M. S. / Museum. / B. M. 1955-354. [w, p]”; 1 ♀ (BMNH), “B.N. BORNEO. / Mt. Kinabalu, / Kiau, 3,000 ft. [p] / 2. 5. [h] 1929. [pink label, p] // Ex F. M. S. / Museum. / B. M. 1955-354. [w, p]”; 1 ♀ (BMNH), “SARAWAK: / C. J. Brooks. / B. M. 1928-193 [w, p]”; 1 ♀ (BMNH), “SARAWAK. / Mt. Matang. [p] / 1500 ft / 30.V [h] 1939 / N. C. E. Miller. [w, p] // Brit. Mus. / 1954-380. [w, p]”; 1 ♂ 1 ♀ (BMNH), “BORNEO, Sabah, ca. 700m / Crocker Range, IV.2013 / Kota Kinabalu→Tambunan / Kipandi Butterfly park, / N5°52 ' 23.5 '' E116°15 ' 0 6.6 '' / General collecting [w, p] // BMNH(E) 2013-58 / M.V.L.Barclay, B.H. Garner / H. Mendel & A. Giusti [w, p]”; 2 ♀ (BMNH), “BORNEO, Sabah, 1240m / Crocker Range, IV.2013 / Kota Kinabalu→Tambunan / N5°51 ' 33.7 '' E116°17 ' 24.1 '' / General collecting [w, p] // M.V.L.Barclay, B.H. Garner / H. Mendel & A. Giusti / BMNH(E) 2013-58 [w, p]”; 2 ♀ (BMNH), “BORNEO, Sabah, 1240m / Crocker Range, IV.2013 / Kota Kinabalu→Tambunan / N5°51 ' 33.7 '' E116°17 ' 24.1 '' / At light sheet [w, p] // M.V.L.Barclay, B.H. Garner / H. Mendel & A. Giusti / BMNH(E) 2013-58 [w, p]”; 18 ♀ (BMNH), “BORNEO, Sabah, 1240m / Crocker Range, IV.2013 / Kota Kinabalu→Tambunan / N5°51 ' 33.7 '' E116°17 ' 24.1 '' / At light sheet [w, p] // BMNH(E) 2013- 58 / B.H. Garner, M.V.L.Barclay / H. Mendel & A. Giusti [w, p]”; 1 ♀ (BMNH), “Borneo / Sarawak / Trusan / Jan: 1915 / Frof: H. W. Smith [h, w] // Ex F. M. S. / Museum. / B. M. 1955-354. [w, p]”; 1 ♀ (BMNH), “Baram / Sarawak [p] / 12.9.19 [h] 20 / J. C. / Moulton [w, p] // Ex F. M. S. / Museum. / B. M. 1955-354. [w, p]”; 1 ♀ (BMNH), “SARAWAK. / Mt. Matang. [p] / 1500 ft / 8.vi. [h] 1939. / N. C. E. Miller. [w, p] // Brit. Mus. / 1954- 380. [w, p]”; 1 ♂ (BMNH), “Mt. Matang, / W. Sarawak. / XII.̕ 13.-I. ̕14. / G. E. Bryant. [w, p] // G. E. Bryant Coll. / 1919-147 [w, p]”; 1 ♀ (BMNH), “Sheet (light). [w, p] // SARAWAK: / Mt. Dulit, / 4.000 ft. / Moss forest. / 24.x.1932 [w, p] // Oxford Univ. Exp. / B. M. Hobby & / A. W. Moore. / B. M. 1933-254. [w, p]”; 1 ♂ (BMNH), “B.N. BORNEO. / Mt. Kinabalu, / Kenokok, [p] / 3,000 [h] ft. [p] / 26 [h] Apr. 1929. [pink label, p] // Ex F. M. S. / Museum. / B. M. 1955-354. [w, p]”; 1 ♂ (BMNH), “Long Mujan [h] / Baram River / Sarawak. [p] / 5.10.19 [h] 20 / J. C. Moulton [w, p] // Ex F. M. S. / Museum. / B. M. 1955-354. [w, p]”; 2 ♂ (BMNH), “Mt. Matang, / W Sarawak. / Dec. 1913. / G. E. Bryant. [w, p] // G. E. Bryant Coll. / 1919-147 [w, p]”; 1 ♀ (BMNH), “FED. MALAY STATES / 1909. / C. J. Brooks. / B. M. 1931-570 [w, p]”; 1 ♀ (BMNH), “Sanga Sanga, / Moorjawa, / E. Borneo. / H. D. Jensen. / 1906-45. [w, p]”; 1 ♀ (BMNH), “SARAWAK: Gungong / Mulu Nat. Park / R.G.S. Exped. 1977-8 / J. D. Holloway et al. / B. M. 1978-206 [w, p] // Site 24. April / W. Melinau Gorge / 270m. 430558 / FEG 4. Limestone / forest, Acl-understorey [w, p] // at / light [w, p]”; 4 ♂ 8 ♀ (NHMB), “Borneo, Sabah, Km 53 / road KK-Tambunan / E slope Gn. Emas, 700 m / 1.–5. Apr 2000, Bolm lgt. [w, p]”; 2 ♂ 7 ♀ (UKM), “SARAWAK: Lanjak / Entimau / 28–29 Feb. 1992 / Zaidi [w, p]”; 3 ♀ (UKM), “SABAH: Tawau / Tibow / 5–10 April 2000 / G. F. Barroga [w, p]”; 2 ♂ 3 ♀ (UKM), “SABAH: Tawau / Tibow / 5–10 April 2000 / Salleh, Ismail & Ruslan [w, p]”; 1 ♂ (UKM), “SABAH, Sipitang / Mondolong ([p] 800 [h] m) [p] / 21 [h] –VIII, 1 988 [p] / Azizah [h] leg. [w, p]”; 1 ♂ (UKM), “SABAH: Lembah Danum / 27–31 Ogos [p] 1991 [h] / Salleh, Zaidi, Mail & Lan [w, p]”; 1 ♂ (UKM), “SABAH: Taman / Kinabalu, Poring / 9–12 Dis. 1990 / Zaidi, Ismail & Ruslan [w, p] // GALERUCINAE [p] / Coeligetes / unicolor Jacoby [h] / det. Mohamedsaid 199 [p] 3 [w, h]”; 1 ♂ (UKM), “SARAWAK: / Bintulu / 17–18 Jan. 1992 / Salleh & Zaidi [w, p] // GALERUCINAE [p] / Coeligetes / unicolor Jacoby [h] / det. Mohamedsaid 199 [p] 3 [w, h]”; 1 ♂ (UKM), “SARAWAK: / Bintulu / 17–18 Jan. 1992 / Salleh & Zaidi [w, p]”; 2 ♀ (UKM), “SABAH: Lembah / Danum / 5–8 Dis. 1992 / Ismail, Yusof, Razali [w, p]”; 1 ♀ (UKM), “SABAH: Lembah / Danum / 22–25 Ogos 1992 / Ismail, Yusof, Sham [w, p]”; 1 ♀ (UKM), “SABAH: Lembah / Danum / 17– 20 April 92 / Ismail, Yusof, Razali [w, p] // GALERUCINAE [p] / Coeligetes / unicolor Jacoby [h] / det. Mohamedsaid 199 [p] 4 [w, p]”; 1 ♂ (UKM), “SARAWAK: Bario / Kg. Alur Palang (H. Primer) / 12 April 1995 / Ismail & Ruslan [w, p]”; 1 ♂ (UKM), “SABAH: Pulau Banggi / 8–12 Mei 1996 / Salleh, Zaidi, Ismail & Sham [w, p] // GALERUCINAE [p] / Coeligetes / unicolor Jacoby [h] / det. Mohamedsaid 199 [p] 6 [w, p]”; 1 ♂ (UKM), “SABAH: Pulau Banggi / 8–12 Mei 1996 / Salleh, Zaidi, Ismail & Sham [w, p]”; 1 ♀ (UKM), “SARAWAK: Taman / Negara Lambir / 1–4 Sept. 1993 / Ismail, Sham, Zabidi [w, p] // GALERUCINAE [p] / Coeligetes / unicolor Jacoby [h] / det. Mohamedsaid 199 [p] 3 [w, h]”; 1 ♂ (UKM), “SABAH: Gunung / Kinabalu, Sayap / 3–8 Jun 1992 / Zaidi, Ismail, Ruslan [w, p] // GALERUCINAE [p] / Coeligetes / unicolor Jacoby [h] / det. Mohamedsaid 199 [p] 3 [w, h]”; 1 ♀ (UKM), “SABAH: Gunung / Kinabalu, Sayap / 3–8 Jun 1992 / Zaidi, Ismail, Ruslan [w, p]”; 1 ♀ (RBCN), “Sabah, Malaysia / Mount Kinabalu / Sayap [p] / 15.6.93 [h] / leg. Hoffmann [w, p] / / Coelligetes / unicolor Jacoby [h] / R. Beenen det. [p]´95 [w, h] // Collection / Ron Beenen [y, p]”; 1 ♂ (UKM), “SABAH: Taman / Kinabalu, HQ / 2–4 Mei 1998 / Zaidi, Ismail, Ruslan [w, p]”; 1 ♂ (BMNH), “Mt. Matang, / W. Sarawak, / G. E. Bryant [p] / xii.1913 [w, h] // G. Bryant Coll. / 1919-147 [w, p]”; 1 ♂ (BMNH), “Mt. Matang, / W. Sarawak. / Jan. 1914. / G. E. Bryant [w, p] // G. Bryant Coll. / 1919-147 [w, p]”; 1 ♀ (BMNH), “Borneo [w, h] // 619. [w, h] // Andrewes / Bequest. / B. M. 1922-221 [w, p]”; 1 ♂ 1 ♀ (BMNH), “BRUNEI: x-1992 / Temburong District / Ridge NE of Kuala / Belagong, 300m. [w, p] // 125W MV Light Trap / J H Martin coll. / BMNH(E) 1992- 172 [w, p]”; 1 ♀ (BMNH), “Borneo / Kinabalu [w, h]”; 1 ♀ (BMNH), “Mt. Matang, / W. Sarawak. / I.–II.1914. / G. E. Bryant [w, p] // G. Bryant Coll. / 1919-147 [w, p]”; 2 ♀ (BMNH), “Mt. Matang, / W. Sarawak, / G. E. Bryant [p] / i.1914 [w, h] // G. Bryant Coll. / 1919-147 [w, p]”; 1 ♀ (BMNH), “Quop, / W. Sarawak, / G. E. Bryant [p] / iv.1914 [w, h] // G. Bryant Coll. / 1919-147 [w, p]”; 1 ♀ (BMNH), “Quop, / W. Sarawak, / II–III.1914. / G. E. Bryant. [w, p] // G. Bryant Coll. / 1919-147 [w, p]”; 3 ♂ (BMNH), “Doherty [w, p] // Borneo / Pengaron [w, p] // Fry Coll. / 1905.100 [w, p]”; 1 ♂ (BMNH), “59471 [w, h] // Doherty [w, p] // Borneo / Pengaron [w, p] // Fry Coll. / 1905.100 [w, p]”; 1 ♀ (BMNH), “Borneo [w, h] // Matang [h] / Dec. / 189 [p] 8 [w, p] // Jacoby Coll. / 1909-28 a. [w, p]”; 2 ♀ (BMNH), “Whitehead [w, p] // Borneo / Kina Balu [w, p] // Fry Coll. / 1905.100 [w, p]”; 7 ♀ (TARI), “Malaysia: Sabah / Trusmadi / 27.IX.2014. leg. Y.-T. Wang [w, p]”; 1 ♂ 10 ♀ (TARI), same data, but 28.IX.2014; 3 ♂ 25 ♀ (TARI), same data, but 29.IX.2014; 1 ♂ 4 ♀ (TARI), same data, but 01.X.2014; 4 ♀ (TARI), same data, but 02.X.2014; 1 ♀ (JVCM), “BORNEO, SABACH [sic!] W. / route RANAU—TAMBUNAN / 2.2000 / Lgt. Snizek [w, p] // ex coll. Daccordi [w, h]”. The specimens are provided with one additional printed red label: “HOLOTYPUS, [or PARATYPUS, respectively], / Coeligetes / howardi sp. nov., / J. Bezděk det., 2015”.

Description. Measurements. Males: 7.3–9.4 mm (holotype 8.8 mm), females: 9.5–13.0 mm.

Body almost completely yellowish brown, except black apices of mandibles.

Male (holotype, Fig. 26 ). Labrum transverse, 2.15 times as wide as long, anterior margin slightly rounded, anterior angles widely rounded, surface subopaque, with 8 pale setae in transverse row. Anterior part of head impunctate, lustrous, convex, with several long pale setae laterally along anterior margin and laterally along antennal insertions. Interantennal space 0.9 times wide as transverse diameter of antennal socket. Eyes large, interocular space narrow, 0.9 times as wide as transverse diameter of eye. Frontal tubercles subpentagonal, transverse, with anterior tips divergent, surface impunctate, dull. Vertex distinctly impressed behind frontal tubercles, surface glabrous, covered with fine punctures, medially behind frontal tubercles with indistinct narrow impunctate line. One setigerous pore with long pale seta placed behind inner posterior eye margin. Antennae thin, 0.85 times as long body, length ratio of antennomeres equals 15-5-12-17-17 -17-17-16-15-14-19.

Pronotum transverse, 2.11 times as broad as long, lustrous, moderately convex, widest in posterior half, sparsely covered with extremely fine punctures. Anterior margin widely concave, lateral margins parallel in posterior half, convergent and rounded anteriorly, posterior margin widely rounded. All margins distinctly bordered, border at posterior half of lateral margins slightly wider. Anterior angles swollen, posterior angles rectangular, all angles with setigerous pores bearing long pale seta, additional short setae visible on anterior half of lateral margins. Scutellum subtriangular, with rounded tip, glabrous, covered with indistinct fine punctures.

Elytra 1.83 times as long as wide and 0.75 times as long as body, almost glabrous, with several short setae visible on lateral and apical slopes, covered with dense confused punctures. Humeral calli well developed. Epipleura wide basally, narrowing at anterior third, thin posteriorly and disappearing before apex.

Anterior coxal cavities closed posteriorly. Last ventrite with two narrow incisions reaching ca 1/3 of ventrite lenght, median lobe with elongate oval impression, penultimate ventrite with small semicircular median concavity ( Fig. 39 ). Protarsomere I narrowly elongate subtriangular, 1.60 times as long as wide, protarsomere II triangular, length ratio of protarsomeres I–IV equals 8-6-7-10. Metatarsomere I long and narrow, length ratio of metatarsomeres I–IV equals 15-7-7-10. Claws appendiculate ( Fig. 36 ).

Aedeagus small, apical half subtriangular, ventrally with three elongate impressions. Dorsal process bulbous, apically with two flat processes, dorsally with large tuft of setae ( Fig. 35 ).

Female ( Fig. 27 ). Eyes smaller, interocular space wider, 1.66 time as wide as transverse diameter of eye. Antennae filiform, slightly thinner and shorter than in male, 0.73 times as long as body. Last abdominal ventrite with deep U-shaped incision in middle of posterior margin ( Fig. 37 ). Pygidium transverse, with posterior margin almost straight, tergite VIII trapezoidal with posterior margin straight ( Fig. 38 ). Sternite VIII rhomboidal, short pale setae cummulated in posterior half, tignum short, about 0.75 time as long as sternite VIII ( Fig. 40 ). Vaginal sclerite elongate semicircular with two hook-like darker lateral patches ( Fig. 41 ). Spermatheca as in Fig. 42 .

Differential diagnosis. In habitus, Coeligetes howardi sp. nov. is very similar to C. unicolor from Java, but differs in having aedeagus wide with apex subtriangular, ventrally with three longitudinal impressions subapically, and dorsal proces extremely bulbous with two flat processes anteriorly ( Fig. 35 ). In C. unicolor the aedeagus is narrow, gradually narrowing towards apex, without any ventral impressions, and dorsal proces is not extremely bulbous, but with two small teeth laterally ( Fig. 28 ). Penultimate abdominal ventrite in males of C. howardi sp. nov. has a shallow semicircular impression in the middle of its basal half, while it is longitudinally impressed with two sharp teeth in C. unicolor ( Figs 29 , 39 ). Females of both species can be distinguished by the shape of the last abdominal ventrite (with a deep U-shaped incision in C. howardi sp. nov., widely shallowly concave in C. unicolor ) ( Figs 30 , 37 ), by setation on rhomboidal sternite VIII (setae in apical half in C. howardi sp. nov., in apical quarter only in C. unicolor ) ( Figs 34 , 40 ), and by the structure of vaginal sclerite (with two longitudinal stripes in C. howardi sp. nov., darkened only in apices in C. unicolor ) ( Figs 32 , 41 ).

Distribution. Malaysia (Sabah, Sarawak), Indonesia (Borneo), Brunei. The data of C. unicolor from Borneo ( Mohamedsaid 1994, 1997a, b, 2000) refer to C. howardi sp. nov. (voucher specimens examined).

Etymology. Dedicated to Howard Mendel (formerly BMNH) who collected part of the type series of the newly described species.

Coeligetoides gen. nov.

Type species. Coeligetoides trifurcatus sp. nov., by present designation.

Description. Body elongate, yellowish brown, dorsum glabrous, shiny.

Labrum transverse, with concave anterior margin. Anterior part of head flat, posterior tip relatively wide. Interantennal space wide, 1.70 times as wide as transverse diameter of antennal socket. Frontal tubercles subpentagonal, transverse. Vertex distinctly impressed behind frontal tubercles, with median impressed line. Antennae filiform, slightly shorter than body.

Pronotum transverse, 1.85 times as broad as long, covered with double fine punctation. Anterior margin slightly concave, lateral margins moderately rounded, posterior margin widely rounded. Anterior margin with indistinct border, lateral and posterior margins bordered. All angles with setigerous pore bearing long pale seta. Scutellum with very widely rounded tip.

Elytra with well developed humeral calli. Surface very densely covered with small confused punctures, almost glabrous, with several short setae visible on lateral and apical slopes. Epipleura wide basally, gradually narrowing posteriorly and disappearing before apex. Macropterous.

Posterior margin of anterior coxal cavities open. Last ventrite with elongate oval impression in median lobe wide, penultimate ventrite with small semicircular incision in middle of posterior margin. Claws bifid.

Aedeagus wide and flat with dorsal process thin and trifurcate. Vaginal sclerite absent, spermatheca with poorly delimited nodulus.

Differential diagnosis. Coeligetoides gen. nov. shares unspined tibiae with Coeligetes but differs in posterior margin of anterior coxal cavities open (closed in Coeligetes ), very unusual wide and flat aedeagus with dorsal process thin and trifurcate, anterior part of head flat with wide posterior tip (elevated and convex with posterior tip narrower in Coeligetes ), claws bifid (claws with small transverse appendix in Coeligetes ), labrum with shallowly concave anterior margin (rounded or nearly straight in Coeligetes ), female last ventrite with entire and regularly rounded posterior margin (incised in Coeligetes ), vaginal sclerite absent (present in Coeligetes ), spermatheca with poorly delimited nodulus (nodulus well developed in Coeligetes ).

The posterior margin of anterior coxal cavities open is shared with representatives of strictly continental Liroetis . However, Coeligetoides gen. nov. can be distiguished from Liroetis by the structure of aedeagus ( Fig. 45 , for aedeagi of Liroetis see e.g. Jiang 1988), claws bifid (appendiculate in Liroetis ), anterior part of head flat with wide posterior tip (elevated and convex with posterior tip narrower in Liroetis ), spermatheca with poorly delimited nodulus (nodulus well developed in Liroetis ), hind tibia in male without spine (with spine in L. aeneipennis Weise, 1889 , type species of Liroetis ).

The genus Luperogala differs in the presence of a large long sword-like process from posterior margin of abdominal ventrite IV directed posteriorly, which is absent in all genera in the generic group including Coeligetoides gen. nov.

The genus Liroetoides can be distinguished from Coeligetoides gen. nov. by closed posterior margin of anterior coxal cavities (open in Coeligetoides gen. nov.), abdominal ventrite V in male transversely concave or with two transverse cavities (with simple median impression in Coeligetoides gen. nov.), long and narrow aedeagus (wide and flat long in Coeligetoides gen. nov.), dorsal process of aedeagus long and narrow (trifurcate in Coeligetoides gen. nov.).

Etymology. Derived from the name of related genus Coeligetes . Gender: masculine.

Figures

FIGURES 24 – 27. Habitus of Coeligetes. 24 — C. unicolor (male, 8.0 mm); 25 — C. unicolor (female, 11.5 mm); 26 — C. howardi sp. nov. (paratype, male, 8.8 mm); 27 — C. howardi sp. nov. (paratype, female, 12.1 mm).

FIGURES 35 – 42. Details of Coeligetes howardi sp. nov. 35 — aedeagus (dorsal, lateral and ventral views); 36 — claws; 37 — female last abdominal ventrite; 38 — female pygidium and tergite VIII; 39 — last two abdominal ventrites of male; 40 — sternite VIII and tignum; 41 — vaginal sclerite; 42 — spermatheca. Scale bars: 0.25 mm for Fig. 36, 0.5 mm for Figs 35 and 40 – 42, 1 mm for Figs 37 – 39.

FIGURES 28 – 34. Details of Coeligetes unicolor. 28 — aedeagus (dorsal, lateral and ventral views); 29 — last two abdominal ventrites of male; 30 — female last abdominal ventrite; 31 — female pygidium and tergite VIII; 32 — vaginal sclerite; 33 — spermatheca; 34 — sternite VIII and tignum. Scale bars: 0.5 mm for Figs 28 and 32 – 34, 1 mm for Figs 29 – 31.

FIGURES 45 – 51. Details of Coeligetoides trifurcatus sp. nov. 45 — aedeagus (dorsal, lateral and ventral views); 46 — claws; 47 — last two abdominal ventrites of male; 48 — female last abdominal ventrite; 49 — female pygidium and tergite VIII; 50 — spermatheca; 51 — sternite VIII and tignum. Scale bars: 0.25 mm for Fig. 46, 0.5 mm for Figs 45, 50 and 51, 1 mm for Figs 47 – 49.