Carya NUTT., 1818

Teodoridis, Vasilis, Kvaček, Zlatko, Radoň, Miroslav, Raprich, Vladislav & Bruch, Angela A., 2020, A New Oligocene Flora From Ludvíkovice Near Děčín (České Středohoří Mts., The Czech Republic), Fossil Imprint 76 (2), pp. 297-314 : 303

Genus Carya NUTT., 1818

Carya fragiliformis (STERNB.) KVAČEK et H.WALTHER in

Walter and Kvaček 2007 / Carya quadrangula (KIRCHH.) J.-F. LEROY, 1955

Pl. 3, Figs 6–9, Pl. 4, Fig. 1

1825 Phyllites fragiliformis STERNB. , p. 42, index iconum, pl. 50, fig. 1.

1936 Caryojuglans quadrangular KIRCHH. , pp. 82–85, text-fig. 13, pl. 12, fig. 36a–l.

1955 Carya quadrangular (KIRCHH.) J.-F.LEROY, pp. 162–163, text-fig. 81.

2007 Carya fragiliformis (STERNB.) KVAČEK et H.WALTHER in Walter and Kvaček , p. 110, text-fig. 6b, pl. 11, figs 1–3, pl. 23, figs 8–10.

M a t e r i a l. Incomplete leaflets and one incomplete endocarp, impressions PA 1627, PA 1644, PA 1659, PA 1660, PA 1682 (fruit).

D e s c r i p t i o n. Detached incomplete leaflets, lanceolate to oblanceolate, 35 to 63 mm long, 12 to 26 mm wide, apex acute to acuminate, base cuneate with fragmentary petiolule up to 5 mm, often asymmetrical, margin irregularly serrate, teeth triangular, sharp, bent admedially, venation craspedodromous, to semicraspedodromous, midvein straight or curved, secondaries opposite in the basal part then alternate, originating at 40° to 60°, tertiary veins percurrent, venation of the higher orders regular polygonal reticulate, areolation well-developed, 3 to 4 sided. Incomplete endocarp widely elliptic in outline, apex acute, upper surface and distinct longitudinal ribs are not visible, base not preserved, 22 mm long and 18 mm wide, thickness of the nutshell 2.8 mm, the primary and secondary septum fragmentary. Seed body (kernel) oval incomplete, 2 lateral symmetric parts preserved.

R e m a r k s. The material corresponds to the holotype of this fossil species based on a single specimen from Žichov ( Kvaček and Straková 1997, Walther and Kvaček 2007 ). The hickory leaves and/or leaflets described as C. fragiliformis or C. serrifolia (GÖPP.) KRÄUSEL are relatively common elements from Oligocene localities in Central Europe, known from e.g., Bechlejovice ( Kvaček and Walther 2004), Suletice-Berand ( Kvaček and Walther 1995), Seifhennersdorf ( Walther and Kvaček 2007 ) and Matrý ( Kvaček et al. 2018). According to Walther and Kvaček (2007: 110) , C. serrifolia , which in most cases was assigned to Oligocene – early Miocene material from Central Europe, may represent a descendant of this lineage in the late Neogene. Koutecký et al. (2019) associated this type of foliage with fruits of C. quadrangula (KIRCHH.) J.-F. LEROY and wood of Eucaryoxylon crystallophorum MÜLL. - STOLL et MÄDEL following the whole plant concept. The fruit find from Ludvíkovice morphologically corresponds to C. quadrangula . On the other hand, some poorly preserved leaf material may show affinity to Engelhardia orsbergensis (P.WESSEL et C.O.WEBER) JÄHNICHEN, MAI et H.WALTHER.