Corythalia tribulosa, Bayer & Höfer & Metzner, 2020

Bayer, Steffen, Höfer, Hubert & Metzner, Heiko, 2020, Revision of the genus Corythalia C. L. Koch, 1850, part 1: Diagnosis and new species from South America (Araneae: Salticidae: Salticinae: Euophryini), Zootaxa 4806 (1), pp. 1-144 : 97-98

Corythalia tribulosa sp. nov.

Figs 47 A–B, 60B, 67A, 70E

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Type material. Holotype: ♂, COLOMBIA: Departamento de Putumayo: El Pepino , about 1000 m a.s.l., 01°03”N, 76°37’50”W, Prof. Dr Norbert Leist leg. 21 Feb. 1973, SMNK-ARA 12954 .

Etymology. The specific name refers to the thorn-like apophyses at the embolus of the male holotype of this species (Latin adjective “tribulosus” meaning “thorny”); adjective.

Diagnosis. Males distinguished from those of all other Corythalia species by the following characters in combination: embolus (E) quite strong, with two conspicuous apophyses centrally ( Figs 47A , 67A ) which are crossing in retrolateral view ( Figs 47B , 70E ) and distally with retrolatero-distal, relatively long and very narrow extension with tip having (disto-) retrolateral orientation ( Figs 47 A–B, 67A) and prolatero-distally with stout, conical and very small process; distal margin of base of E (EB) in ventral view almost reaching distal margin of tegulum (T) (gap <width of RTA in ventral view) ( Figs 47A , 67A ); RTA medium-sized (shorter than width of T), without dorsal serration and with tip distinctly bent ventrally ( Fig. 47B ).

Description. Male: total length 5.6, carapace length 2.5, maximal carapace width 1.9, width of eye rectangle 1.7, opisthosoma length 2.6, opisthosoma width 1.5, fovea length 0.28. EYES: AME 0.58, ALE 0.36, PME 0.08, PLE 0.33, AME–AME 0.03, AME–ALE 0.03, PME–PME 1.46, PME–PLE 0.22, ALE–PLE 0.61, PLE–PLE 1.19, clypeus height at AME 0.29, clypeus height at ALE 0.62. Cheliceral furrow with 2 (one of them slightly smaller, both located directly next to each other) promarginal and 1 retromarginal teeth. SPINATION: palp without spines. Legs: femur I 1500, II 1500 {1600}, III 1500, IV 1600; patella I 1000, II–IV 1010; tibia I 1003 {2004}, II 3013, III–IV 3133; metatarsus I 2014, II 2024, III 3134, IV 4143{4144}. MEASUREMENT OF PALP AND LEGS: palp 2.0 [0.8, 0.3, 0.2, 0.7], I 4.4 [1.5, 0.7, 0.9, 0.8, 0.5], II 4.6 [1.5, 0.7, 1.0, 0.9, 0.5], III 5.9 [1.9, 0.8, 1.3, 1.3, 0.6], IV 6.0 [1.8, 0.8, 1.3, 1.5, 0.6]. LEG FORMULA: 4321. COPULATORY ORGAN: embolus (E) (actual tubular section) moderately long [slightly shorter than width of tegulum (T)], quite broad [centrally (even without apophyses) slightly broader than RTA in ventral view], retrolatero-distally with quite long, light and very narrow extension with tip having (disto-) retrolateral orientation ( Figs 47A , 67A ) and prolatero-distally with stout, conical and very small process, centrally with thorn-shaped apophysis prolaterally and quite prominent and pointed apophysis retrolaterally ( Figs 47A , 67A ); width of embolus base (EB) circle> 1/2, but <2/3 the width of T; EB located centrally to prolatero-centrally at distal part of T; T narrower than cymbium ( Figs 47A , 67A ); sperm duct double-stacked S-shaped, occupying more than 2/3 of T from retrolateral; proximal tegulum lobe not recognisable as such, but T proximally moderately converging and proximal ending (very) broad rounded; cymbium in ventral view distally conically converging, at distalmost section rounded; palpal tibia short, broader than long ( Figs 47 A–B, 67A, 70E) and ventral tibial bump in ventral view broad, distally almost truncated, located in proximal section of prolateral half of palpal tibia and having prolatero-distal orientation; RTA in ventral view quite narrow, medium-sized, with almost distal direction and without dorsal serration ( Figs 47A , 67A ), in retrolateral view RTA distally converging and with tip clearly bent ventrally ( Figs 47B , 70E ). COLOURATION: see genus description for conservative aspects. Carapace red-brown, proximal sections of lateral margins without broad bands of dense, light scale hairs ( Fig. 60B ). Legs brown to red-brown, except for proximalmost articles, patellae and tarsi being lighter ( Fig. 60B ). Opisthosoma like noted in genus description under general dorsal colouration, however central transversal band very narrow (just as narrow as posterior band) and quite even and straight, chevron-like patch in central band missing; posterior band distinctly separated medially ( Fig. 60B ).

Female: unknown.

Remarks. Among the currently known Corythalia species, C. valida , without any doubt, is most similar to C. tribulosa sp. nov. Several crucial structures of the palp are very similar: embolus (E) quite broad and strong, centrally with apophyses (three in C. valida , two in C. tribulosa sp. nov.); similar distal section of E; tegulum without distinguished proximal lobe, but broad rounded proximally; slender RTA with (at least slight) bending ventrally. Consequently, we expect a close relationship between these two species.

Distribution. Currently known only from the type locality in Putumayo, Colombia.

Figures

FIGURE 47. Corythalia tribulosa sp. nov., male holotype from Departamento de Putumayo, Colombia. A–B Left palp (A ventral view; B retrolateral view).

FIGURE 60. Corythalia spp., photographic images of male specimens, dorsal view. Corythalia hamulifera Bayer, sp. nov.: A Holotype from south-eastern part of Ecuador. Corythalia tribulosa sp. nov.: B Holotype from Putumayo, Colombia. Corythalia valida: C Lectotype from Amazonas, Brazil. Corythalia flagrans sp. nov.: D Holotype from Acre, Brazil. Corythalia fragilis sp. nov.: E Paratype SMNK-ARA 02236 from Amazonas, Brazil. Corythalia grata F Lectotype from São Paulo, Brazil. Corythalia vervloeti: G Male M-1 from Borba, Amazonas, Brazil. Corythalia opima: H Male SMF 8437 from La Paz, El Salvador. Corythalia spiralis: I Lectotype from Chiriquí, Panama.

FIGURE 67. Corythalia spp., photographic images of left palps of male specimens, ventral view. Corythalia tribulosa sp. nov.: A Holotype from Putumayo, Colombia. Corythalia valida. B Lectotype from Amazonas, Brazil. Corythalia flagrans sp. nov.: C Holotype from Acre, Brazil. Corythalia grata: D Lectotype from São Paulo, Brazil. Corythalia fragilis sp. nov.: Paratypes E SMNK-ARA 02232, F SMNK-ARA 02236, G SMNK-ARA 02237-1 and H SMNK-ARA 02237-2 from Amazonas, Brazil. Corythalia vervloeti: I Male M-2 from Amazonas and J M-4 from Paraná, Brazil. Corythalia opima: K Lectotype from eastern part of Guatemala. Corythalia spiralis: L Lectotype from Chiriquí, Panama.

FIGURE 70. Corythalia spp., photographic images of left palps of male specimens, retrolateral view. Corythalia hadzji: A Lectotype from Upper Demerara-Berbice, Guyana. Corythalia trochophora Bayer, sp. nov.: B Holotype from Tungurahua, Ecuador. Corythalia lineata Bayer, sp. nov.: C Holotype from East Berbice-Corentyne, Guyana. Corythalia hamulifera Bayer, sp. nov.: D Holotype from south-eastern part of Ecuador. Corythalia tribulosa sp. nov.: E Holotype from Putumayo, Colombia. Corythalia valida: F Lectotype from Amazonas, Brazil. Corythalia flagrans sp. nov.: G Holotype from Acre, Brazil. Corythalia fragilis sp. nov.: H Paratype SMNK-ARA 02236 from Amazonas, Brazil. Corythalia grata: I Lectotype from São Paulo, Brazil. Corythalia vervloeti: J Male M-2 from Amazonas, Brazil. Corythalia opima: K Lectotype from eastern part of Guatemala. Corythalia spiralis: L Lectotype from Chiriquí, Panama.