Corythalia fimbriata ( Peckham & Peckham, 1901 )

Bayer, Steffen, Höfer, Hubert & Metzner, Heiko, 2020, Revision of the genus Corythalia C. L. Koch, 1850, part 1: Diagnosis and new species from South America (Araneae: Salticidae: Salticinae: Euophryini), Zootaxa 4806 (1), pp. 1-144 : 41-43

Corythalia fimbriata ( Peckham & Peckham, 1901)

Figs 4F , 20 A–D, 58A, 65B, 68J

Dynamius fimbriatus Peckham & Peckham 1901: 340 , pl. 25, fig. 10, pl. 26, fig. 12 (description & illustration of ♂). Lectotype ♂ (here designated) from Brazil: Bahia: Chapada, meaning Parque Nacional da Chapada Diamantina , in Peckham & Peck- ham (1901) still listed as Chapoda; G.W. & E.G. Peckham Coll., No. 661 (obtained from H. H. Smith Coll.); MCZ 21310; 1 ♂ paralectotype (here designated) with exactly the same data as for lectotype; MCZ 21310, all type material examined.

Corythalia fimbriata — Petrunkevitch 1911: 616 (transfer from Dynamius to Corythalia ).

Diagnosis. Males distinguished from those of all other Corythalia species by the following characters in combination: embolus (E) very short [just about 1/2 the width of tegulum (T)] and base of E (EB) very small (width of EB circle at most 1/3 the width of T); E without special structures and continuously converging from central to (approximately) pointed distal section ( Figs 20A, 20C & 65B ); T retrolatero-proximally with distinct projecting lobe.

Description. Male (measurements of lectotype first, those of paralectotype in parentheses): total length 7.0 (6.2), carapace length 3.4 (3.1), maximal carapace width 2.5 (2.2), width of eye rectangle 1.9 (1.7), opisthosoma length 3.4 (2.9), opisthosoma width 2.3 (1.9), fovea length 0.31 (0.22). EYES: AME 0.56 (0.51), ALE 0.34 (0.32), PME 0.13 (0.10), PLE 0.32 (0.28), AME–AME 0.06 (0.05), AME–ALE 0.11 (0.08), PME–PME 1.68 (1.48), PME– PLE 0.28 (0.27), ALE–PLE 0.77 (0.73), PLE–PLE 1.43 (1.25), clypeus height at AME 0.38 (0.33), clypeus height at ALE 0.73 (0.69). Cheliceral furrow with 1 promarginal and 1 retromarginal teeth. Cymbium scopula consisting of light hairs in this species minimally more dense and conspicuous ( Figs 3F , 38F ) than in other Corythalia species. SPINATION: palp: no spines. Legs: femur I 1500, II 1600, III 1500, IV 1600; patella I–II 1000, III–IV 1010; tibia I 2004, II 3006 (2004), III 3134 (3133), IV 3134; metatarsus I 2024 (2014), II 2024, III 3134 (4134), IV 4134 (4134{5144}). MEASUREMENT OF PALP AND LEGS: palp 2.6 (2.3) [1.0 (0.9), 0.4, 0.3 (0.2), 0.9 (0.8)], I 5.9 (5.3) [2.0 (1.8), 1.1 (0.9), 1.2 (1.1), 1.0 (0.9), 0.6.], II 6.3 (5.5) [2.1 (1.8), 1.2 (1.0), 1.3 (1.1), 1.1 (1.0), 0.6], III 7.6 (6.7) [2.4 (2.2), 1.2 (1.1.), 1.6 (1.4), 1.6 (1.4), 0.8 (0.6)], IV 7.1 (6.3) [2.2 (1.9), 1.1 (1.0), 1.4 (1.3), 1.6 (1.4), 0.8 (0.7)]. LEG FORMULA: 3421. COPULATORY ORGAN: embolus (E) very short [just slightly longer than 1/2 the width of tegulum (T)], sickle-shaped, arising centro- to prolatero-proximal at embolus base (EB); EB small (width of EB circle at most 1/3 the width of T) ( Figs 20A, 20C & 65B ). T slightly narrower than cymbium, sperm duct double-stacked S-shaped, occupying slightly more than retrolateral 1/2 of tegulum ( Figs 20A, 20C & 65B ). Cymbium (CY) in ventral view conically converging distally, distalmost section still broadly rounded ( Figs 20A, 20C & 65B ). Palpal tibia moderately short (in comparision to most other Corythalia species), just slightly broader than long ( Figs 20 A–D & 65B, 68J) and ventral tibial bump conical, distally pointed and directed distally ( Figs 20A, 20C & 65B ). RTA in ventral view quite slim, in retrolateral view with slightly ventral direction (not straight distal direction) and at distal section blunt converging, in contrast to ventral view dorsal serration in retetrolateral view not recognisable ( Figs 20B, 20D & 68J ). Scopula, like in all Corythalia species, only at distal section of dorsal CY (in C. fimbriata distal 1/4) and not distinctly dense. CY at distal 1/4 lighter than further proximally ( Figs 4F , 68J ). COLOURATION: see genus description for conservative aspects. Carapace dark red-brown ( Fig. 58A ). Legs dark brown to red-brown, except for some articles being lighter (see genus description) ( Fig. 58A ). Opisthosoma like noted in genus description under general dorsal colouration, chevron-like patch in central band at most very inconspicuously recognisable ( Fig. 58A ).

Female: unknown.

Intraspecific variation of male copulatory organs. Lectotype male with minimally shorter embolus than paralectotype and homogeneous distal tegulum margin ( Figs 20A , 65B ). In paralectotype retrolatero-distal part of tegulum slightly bulging out distally ( Fig. 20C ). Central loop of sperm duct in lectotype slightly narrower than in paralectotype. RTA in retrolateral view in lectotype ( Figs 20B , 68J ) slightly more directed ventrally and cymbium in retrolateral view distally slightly truncated whereas in paralectotype ( Fig. 20D ) RTA is slightly less directed ventrally and cymbium distally rather rounded.

Remarks. At arrival at SMNK (Apr. 2015) the male lectotype and the male paralectotype were found among the types of Dynamius parvus (MCZ 22546). They must have been erroneously put there. G. Bodner (Mar. 2000) already recognized that these two males differed from the 5 remaining males (meaning those indeed belonging to D. parvus ) which match the original figures in Peckham & Peckham (1901) for D. parvus . The two males described above, however, definitely match the drawings for D. fimbriatus in Peckham & Peckham (1901) ! Hence, they were separated from the series with the types of D. parvus , but with clear statement from which vial they were taken. The above mentioned explains that after our request the types of D. fimbriatus were not found by L. Leibensperger, the curator of MCZ.

Distribution. Known only from the type locality in Chapada Diamantina, Bahia, Brazil.

Figures

FIGURE 3. Corythalia spp., photographic images of habitus and some somatic characters. Corythalia protensa sp. nov., male holotype from Amazonas, Brazil. A Cheliceral bases and claws, labium and gnathocoxae, ventral view. Corythalia fragilis sp. nov., male paratype SMNK-ARA 02236 from Amazonas, Brazil. B Prosoma, frontal view. Corythalia concinna sp. nov., female holotype from Amazonas, Brazil. C Carapace, lateral view. Corythalia foelixi Bayer, sp. nov., male holotype from Cayenne, French Guiana. D Dorsal view. Corythalia valida, male lectotype from Amazonas, Brazil. E Ventral view. Corythalia drepane sp. nov., female paratype SMNK-ARA 13423 from Amazonas, Brazil. F Labium and gnathocoxae, ventral view. Arrows in A indicate very small and slim teeth on retromargins of cheliceral furrows. Arrows in B indicate small lobes on medio-distal section of cheliceral base bearing minimally curved hairs.

FIGURE 4. Corythalia spp., photographic images of habitus, somatic characters and male copulatory organs. Corythalia flagrans sp. nov., male holotype from Acre, Brazil. A Cheliceral bases and claws, labium and gnathocoxae, ventral view; Corythalia luctuosa. B Ventral view, female from St. Laurent du Maroni, French Guiana; D Left palp, prolateral view, male L-1 from St. Laurent du Maroni, French Guiana. Corythalia longiducta sp. nov., female SMNK-ARA 02858 from Amazonas, Brazil. C Prosoma, ventral view, with focus on sternum. Corythalia foelixi Bayer, sp. nov., female from Cayenne, French Guiana. E Spinnerets, ventral view. Corythalia fimbriata, male lectotype from Bahia, Brazil. F Left palp, dorsal view.

FIGURE 20. Corythalia fimbriata, specimens from Bahia, Brazil.A–B Male lectotype, left palp (A ventral view; B retrolateral view). C–D Male paralectotype, left palp (C ventral view; D retrolateral view).

FIGURE 38. Corythalia metallica, male holotype from Saint Vincent Island (Lesser Antilles). A–B left palp (A ventral view; B retrolateral view).

FIGURE 58. Corythalia spp., photographic images of male specimens, dorsal view. Corythalia fimbriata: A Holotype from Bahia, Brazil. Corythalia argentinensis: B Holotype from Misiones, Argentina. Corythalia luctuosa: C Lectotype from Saint Laurent du Maroni, French Guiana. Corythalia cincta: D Holotype from Presidente Hayes, Paraguay. Corythalia scutellaris Bayer, sp. nov.: E Holotype from south-eastern part of Ecuador. Corythalia waleckii: F Syntype MIZ 225794, Cayenne, French Guiana. Corythalia tropica: G Holotype from Falcon, Venezuela. Corythalia foelixi Bayer, sp. nov.: H Holotype from Cayenne, French Guiana. Corythalia fulgipedia: I Male AMNH-IZC 00327930 from Aragua, Venezuela.

FIGURE 65. Corythalia spp., photographic images of left palps of male specimens, ventral view. Corythalia gasnieri sp. nov.:A Holotype from Amazonas, Brazil. Corythalia fimbriata: B Lectotype from Bahia, Brazil. Corythalia argentinensis: C Holotype from Misiones, Argentina. Corythalia luctuosa: D Lectotype from St. Laurent du Maroni, French Guiana. Corythalia cincta: E Holotype and F Male NHMB 1217a (holotype of Taeoma barbipes) from Presidente Hayes, Paraguay. Corythalia scutellaris Bayer, sp. nov.: G Holotype from south-eastern part of Ecuador. Corythalia waleckii: H Syntype MIZ 225794 from Cayenne, French Guiana. Corythalia tropica: I Holotype from Falcon, Venezuela. Corythalia foelixi Bayer, sp. nov.: J Holotype and K Paratype from Cayenne, French Guiana. Corythalia fulgipedia: L Male AMNH-IZC 00327822-1 from Aragua, Venezuela.

FIGURE 68. Corythalia spp., photographic images of left palps of male specimens, retrolateral view. Corythalia latipes: A Neotype from Bahia, Brazil. Corythalia conferta sp. nov.: B Paratype M-1, IBSP 131451, from São Paulo, Brazil. Corythalia drepane sp. nov.: C Holotype from Amazonas, Brazil. Corythalia antepagmenti sp. nov.: D Paratype from Acre, Brazil. Corythalia ricti Bayer, sp. nov.: E Holotype from East Berbice-Corentyne, Guyana. Corythalia parva: F Lectotype from Brazil. Corythalia protensa sp. nov.: G Holotype from Amazonas, Brazil. Corythalia placata: H Male MCZ 20545-II (Paralectotype of Dynamius blandus) from Trinidad. Corythalia gasnieri sp. nov.: I Holotype from Amazonas, Brazil. Corythalia fimbriata: J Lectotype from Bahia, Brazil. Corythalia argentinensis: K Holotype from Misiones, Argentina. Corythalia luctuosa: L Lectotype from St. Laurent du Maroni, French Guiana.