Corythalia protensa, Bayer & Höfer & Metzner, 2020

Bayer, Steffen, Höfer, Hubert & Metzner, Heiko, 2020, Revision of the genus Corythalia C. L. Koch, 1850, part 1: Diagnosis and new species from South America (Araneae: Salticidae: Salticinae: Euophryini), Zootaxa 4806 (1), pp. 1-144 : 35-36

Corythalia protensa sp. nov.

Figs 3A , 17 A–B, 57G, 64J–K, 68G

urn:lsid:zoobank.org:act:F0335FEA-6B07-40EC-A5A0-418452D4EC6D

Type material. Holotype: ♂, BRAZIL: Amazonas : Manaus, Reserva Ducke, ca. 2°55’50”S, 59°58’30”W, noninundated primary forest; H. Höfer & T. Gasnier leg. 05 Oct. 1992, interim deposition SMNK-ARA 02496 , final deposition INPA . Paratype: ♂, with exactly the same data as for holotype, SMNK-ARA 02496 .

Etymology. The specific name refers to the retrolateral tibial apophysis of the male holotype resembling a hand with a long index finger stretched out (Latin “protensa” means “stretched out”, “sprawled out”); adjective.

Diagnosis. Males distinguished from those of all other Corythalia species by the following characters in combination: embolus (E) with very broad proximal section (more than 8x broader than subdistal section), short (approximately as long as width of embolus base) and distally pointed ( Figs 17A , 64 J–K); RTA in retrolateral view very broad and distinctly digitate at ventro-distal section (digitate section with less than 1/5 the width of central section, Figs 17B , 68G ); tegulum (T) (retrolatero-) proximally with distinct projecting lobe (PTL), the latter with flat and blunt bulge prolaterally ( Figs 17A , 64 J–K).

Description. Male: total length 4.7, carapace length 2.1, maximal carapace width 1.6, width of eye rectangle 1.3, opisthosoma length 2.2, opisthosoma width 1.4, fovea length 0.13. EYES: AME 0.44, ALE 0.24, PME 0.07, PLE 0.20, AME–AME 0.03, AME–ALE 0.04, PME–PME 1.09, PME–PLE 0.19, ALE–PLE 0.55, PLE–PLE 0.93, clypeus height at AME 0.17, clypeus height at ALE 0.47. Cheliceral furrow with 1 promarginal and 1 retromarginal teeth. SPINATION: palp: no spines. Legs: femur I 1500, II 1600, III 2600, IV 1600; patella I–II 1000, III–IV 1010; tibia I 2005 {2004}, II 3004{3003}, III–IV 3133; metatarsus I 2014, II 2024 {2013}, III 3134, IV 4144. MEASURE- MENT OF PALP AND LEGS: palp 1.6 [0.6, 0.3, 0.1, 0.6], I 3.4 [1.1, 0.6, 0.7, 0.6, 0.4], II 3.5 [1.1, 0.7, 0.7, 0.6, 0.4], III 4.0 [1.4, 0.6, 0.8, 0.8, 0.4], IV 4.2 [1.3, 0.6, 0.8, 1.0, 0.5]. LEG FORMULA: 4321. COPULATORY OR- GAN: embolus (E) short, basal section very broad, its arising point centro-proximally at embolus base (EB); EB not exactly round, EB circle less than half as broad as tegulum (T) ( Figs 17A , 64 J–K). T slightly narrower than cymbium and proximally of EB with flat triangular lobe directed distally, sperm duct double-stacked S-shaped with quite wide loops (distal loop even slightly wider than proximal), occupying retrolateral 2/3 of T ( Figs 17A , 64 J–K). Cymbium distally in ventral ( Figs 17A , 64 J–K) and lateral view ( Figs 17B , 68G ) slightly truncated. Palpal tibia very short, clearly broader than long ( Figs 17 A–B, 64J–K, 68G) and ventral tibial bump conical and not very conspicuous ( Figs 17A , 64 J–K). RTA broad, slightly directed ventrally and with conspicuous digitate protrusion ventro-distally ( Figs 17B , 68G ) and in dorso-distal section with rounded, finely serrated lobe. COLOURATION: see genus description for conservative aspects. Carapace dark brown to dark red-brown ( Fig. 57G ). Legs dark brown to red-brown, except for some articles being lighter (see genus description) ( Fig. 57G ). Opisthosoma like noted in genus description un- der general dorsal colouration, except for chevron-like patch in central band missing ( Fig. 57G ).

Female: unknown.

Intraspecific variation of male copulatory organs. Flat, triangular lobe proximally of embolus base more distinctly developed in holotype ( Figs 17A , 64J ). Retrolatero-proximal tegulum lobe slightly broader in paratype ( Fig. 64K ). Flat and blunt bulge prolaterally on proximal tegulum lobe in paratype ( Fig. 64K ) larger than in holotype ( Figs 17A , 64J ).

Remarks. This species shows hardly similarities to any other Corythalia species. The shape of tegulum is a little bit reminiscent of that of C. drepane sp. nov. However, most other characters are clearly different, e.g. embolus, RTA or spermduct-course.

Distribution. Known only from the type locality in Central Amazonia, Brazil.

Figures

FIGURE 3. Corythalia spp., photographic images of habitus and some somatic characters. Corythalia protensa sp. nov., male holotype from Amazonas, Brazil. A Cheliceral bases and claws, labium and gnathocoxae, ventral view. Corythalia fragilis sp. nov., male paratype SMNK-ARA 02236 from Amazonas, Brazil. B Prosoma, frontal view. Corythalia concinna sp. nov., female holotype from Amazonas, Brazil. C Carapace, lateral view. Corythalia foelixi Bayer, sp. nov., male holotype from Cayenne, French Guiana. D Dorsal view. Corythalia valida, male lectotype from Amazonas, Brazil. E Ventral view. Corythalia drepane sp. nov., female paratype SMNK-ARA 13423 from Amazonas, Brazil. F Labium and gnathocoxae, ventral view. Arrows in A indicate very small and slim teeth on retromargins of cheliceral furrows. Arrows in B indicate small lobes on medio-distal section of cheliceral base bearing minimally curved hairs.

FIGURE 17. Corythalia protensa sp. nov., male holotype from Amazonas, Brazil.A–B left palp (A ventral view; B retrolateral view).

FIGURE 57. Corythalia spp., photographic images of male specimens, dorsal view. Corythalia latipes: A Neotype from Bahia, Brazil. Corythalia conferta sp. nov.: B Paratype M-1, IBSP 131451, from São Paulo, Brazil. Corythalia drepane sp. nov.: C Holotype from Amazonas, Brazil. Corythalia antepagmenti sp. nov.: D Paratype from Acre, Brazil. Corythalia ricti Bayer, sp. nov.: E Holotype from East Berbice-Corentyne, Guyana. Corythalia parva: F Lectotype from Brazil. Corythalia protensa sp. nov.: G Holotype from Amazonas, Brazil. Corythalia placata: H Male MCZ 20545-II (Paralectotype of Dynamius blandus) from Trinidad. Corythalia gasnieri sp. nov.: I Holotype from Amazonas, Brazil.

FIGURE 64. Corythalia spp., photographic images of left palps of male specimens, ventral view. Corythalia latipes: A Neotype from Bahia, Brazil. Corythalia conferta sp. nov.: Paratypes from São Paulo (B M-9, IBSP 33503; C M-4, IBSP 53033) and Mato Grosso do Sul (D M-6, IBSP 53374), Brazil. Corythalia drepane sp. nov.: E Holotype and F Paratype SMNK-ARA 02326 (in this specimen bulb expanded; thus basal and distal haematodocha recognisable) from Amazonas, Brazil. Corythalia antepagmenti sp. nov.: G Paratype from Acre, Brazil. Corythalia ricti Bayer, sp. nov.: H Holotype from East Berbice-Corentyne, Guyana. Corythalia parva: I Lectotype from Brazil. Corythalia protensa sp. nov.: J Holotype and K paratype from Amazonas, Brazil. Corythalia placata: L Male MCZ 20545-II (Paralectotype of Dynamius blandus) from Trinidad.

FIGURE 68. Corythalia spp., photographic images of left palps of male specimens, retrolateral view. Corythalia latipes: A Neotype from Bahia, Brazil. Corythalia conferta sp. nov.: B Paratype M-1, IBSP 131451, from São Paulo, Brazil. Corythalia drepane sp. nov.: C Holotype from Amazonas, Brazil. Corythalia antepagmenti sp. nov.: D Paratype from Acre, Brazil. Corythalia ricti Bayer, sp. nov.: E Holotype from East Berbice-Corentyne, Guyana. Corythalia parva: F Lectotype from Brazil. Corythalia protensa sp. nov.: G Holotype from Amazonas, Brazil. Corythalia placata: H Male MCZ 20545-II (Paralectotype of Dynamius blandus) from Trinidad. Corythalia gasnieri sp. nov.: I Holotype from Amazonas, Brazil. Corythalia fimbriata: J Lectotype from Bahia, Brazil. Corythalia argentinensis: K Holotype from Misiones, Argentina. Corythalia luctuosa: L Lectotype from St. Laurent du Maroni, French Guiana.