Sagola sheldoni Park and Carlton

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 138

Sagola sheldoni Park and Carlton , new species

( Figs. 68k , 69k , 70k , 72 )

Type Material. Holotype. NEW ZEALAND: Northland: 1♂ ( NZAC), aedeagus dissected and mounted in balsam on a clear plastic card “ NEW ZEALAND ND Ruakaka 22 Jan 1977 C.F. Butcher ”, “ Pit trap in pasture”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola sheldoni Park and Carlton 2013 ” . Paratype (1 female). NEW ZEALAND: Northland : 1♀ ( NZAC), Waipoua SF, 24 XI 1980, G. Kuschel, mats on roadside .

Etymology. This species is named for one of the enthusiastic supporters of this study, Frederick H. Sheldon. a member of the first author’ s PhD advisory committee who provided valuable advice during this study, a major component of his dissertation.

Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.4–2.6 mm; temporal lobe of male head projecting and exceeding eye, with flattened ventral surface; ventral surface of male head with triangular setose process behind mouthparts; elytra subquadrate; shape of antennomeres and genitalia unique to species; only known from Northland.

Description. Length 2.4–2.6 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler ( Fig. 68k ). Head: Male head triangular, widest across temples ( Fig. 69k ). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2–5 longer than wide, 6–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye large and prominent, approximately one-half length of temple. Temporal lobe of male head projecting and exceeding eye, with flattened ventral surface ( Fig. 69k ). Ventral surface of male head with triangular setose process behind mouthparts ( Fig. 69k ). Thorax: Prosternum as long as wide, widest at one-third length. Elytra subquadrate ( Fig. 68k ). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Male tergite IV with pair of small round patches of microtrichia reaching middle, patches absent in female. Aedeagus: Median lobe of genitalia widest at one-third length, with long rectangular apical lobe ( Fig. 70k ). Phallobase of median lobe symmetrical and rounded ( Fig. 70k ). Parameres symmetrical and slender, with sparse setae from apex to midpoint ( Fig. 70k ).

Distribution. Northland ( Fig. 72 : white circles).

Habitat. Specimens of this species were collected using pitfall traps in a pasture or by sifting mats on roadsides.

Figures

Fig. 68. Habitus of Sagola species in the insignis species-group, dorsal views. a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 1 mm.

Fig. 69. Ventral views of male heads of Sagola species in the insignis species-group. a) Sagola insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 0.1 mm.

Fig. 70. Sagola species in the insignis species-group. Aedeagi, dorsal views: a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Male heads, dorsal surface: y) S. bipunctata, z) S. eminen. Scale bars = 0.1 mm.

Fig. 72. Known collection localities of New Zealand Sagola species. S. pertinax: black circles; S. monticola: triangles; S. laticeps: black squares; S. castanea: stars; S. sheldoni: white circles; S. mercuryensis: white squares.