Sagola terricola Broun, 1886, 2013

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 11-12

Sagola terricola Broun, 1886

( Figs. 3b , 4b, 4t –u , 5 )

Sagola terricola Broun, 1886: 832 . Raffray 1893: 34, 1904: 498, 1911: 6, 1924: 231; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 244.

Type Material. Lectotype. NEW ZEALAND: Auckland: 1♂ ( BMNH), glued on rectangular card, “Type” [red label, printed]; “1480. ♂.” [white label, handwritten]; “Waitakerei” [red label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola terricola ” [white label, handwritten]; “ LECTOTYPE Sagola terricola Broun Desig. Park and Carlton 2013 ” [white label, printed]. Paralectotype (1 male). NEW ZEALAND: Auckland: 1♂ ( BMNH), glued on rectangular card, “Type” [red label, printed]; “1480. ♂.” [white label, handwritten]; “Waitakerei” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]. The lectotype designation is required because Broun did not explicitly designate a type specimen, and he mentioned that two specimens were examined ( Broun, 1886: 832). This designation will fix the identity of the species and facilitate its recognition by future workers.

Additional Material (n = 6: 5 males; 1 female). NEW ZEALAND: Northland: 3♂♂ (1♂ slidemounted), Waipoua State Forest, Toatoa Track , 270m, 12–15 IV 1080, toatoa-kauri-podocarpbroadlf, A. Newton, M. Thayer, pitfall trap, berl. leaf and log litter, forest floor; 1♂, Waipoua SF, Ricker Tr. , 100m, 11–15 IV 1980, kauri-podocarpbroadleaf, A. Newton, M. Thayer, pitfall trap 619; 1♂, Waipoua SF, Toronui Track , 120m, 26 XI–4 XII 1984, kauri-podocarp-hdwd, A. Newton, M. Thayer 685, FIT, window trap; 1♀, SH12, Waipoua SF, 20 IX 1977, D. W. Helmore, rotten stumps and trees 77/102.

Diagnosis. Specimens of S. terricola can be distinguished from the other species of the hirtalis species-group by the following combination of characters: larger body size, 2.8–3.5 mm; antennomere 1 elongate, approximately 3 times longer than wide; antennomeres 4–5 elongate, approximately 1.5 times longer than wide; posterior frontal fovea elongate; eye small, one-third length of temple; hind wings well-developed; shape of genitalia unique to species.

Redescription. Length 2.8–3.5 mm. Body brown, elytra, legs, and maxillary palpi yellowish brown ( Fig. 3b ). Head: As long as wide, widest across eyes ( Fig. 3b ). Antennomere 1 approximately 3 times longer than wide, 2 subquadrate, 3 longer than wide, 4–5 elongate, 6 longer than wide, 7–10 subquadrate. Frontal sulcus deep and reaching behind eye from end of frontal rostral lobes. Anterior frontal fovea round, posterior frontal fovea elongate. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular ( Fig. 3b ). Hind wings well-developed. Meso- and metathorax trapezoidal, longer than wide ( Fig. 4t ). Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe longer than parameres with bird-head-shape of apical lobe ( Fig. 4b ). Phallobase of median lobe asymmetrical and triangular ( Fig. 4b ). Left paramere with 2 slender branches from base, right paramere longer than left ( Fig. 4b ).

Type Locality. Waitakerei Range , Auckland, New Zealand.

Distribution. Auckland, Northland ( Fig. 5 : triangles).

Habitat. Specimens of this species were collected using pitfall, window, and flight intercept traps or by sifting leaf or log litter in kauri, broadleaf, hardwood, and podocarp forests.

Figures

Fig. 3. Habitus of Sagola species in the hirtalis species-group, dorsal views. a) Sagola hirtalis, b) S. terricola, c) S. convexa, d) S. coromandelensis, e) S. northlandensis, f) S. taranakiensis, g) S. kuscheli, h) S. newtoni, i) S. thorpei, j) S. dugdalei, k) S. thayerae, l) S. hunuaensis, m) S. waikatoensis, n) S. gisbornensis, o) S. weiri, p) S. brookesi. Scale bars = 1 mm.

Fig. 4. Sagola species in the hirtalis species-group. Aedeagi, dorsal view: a) S. hirtalis, b) S. terricola, c) S. convexa, d) S. coromandelensis, e) S. northlandensis, f) S. taranakiensis, g) S. kuscheli, h) S. newtoni, i) S. thorpei, j) S. dugdalei, k) S. thayerae, l) S. hunuaensis, m) S. waikatoensis, n) S. gisbornensis, o) S. weiri, p) S. brookesi. S. hirtalis: q) Antenna, r) Head, dorsal view, s) Head, ventral view (arrow indicates transverse depression). S. terricola: t) Meso- and metaventrite, u) Male fore femur (arrow indicates semicircular depression). Scale bars = 0.1 mm.

Fig. 5. Known collection localities of New Zealand Sagola species. S. hirtalis: black circles; S. terricola: triangles; S. convexa: black squares; S. coromandelensis: star; S. northlandensis: white circle; S. taranakiensis: white square.