Sagola socia Broun, 1915

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 83-84

Sagola socia Broun, 1915

( Figs. 47a, 47f, 47k , 48 )

Sagola socia Broun, 1915: 281 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 244.

Sagola unicalis Broun, 1917: 376 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 244. New synonymy.

Type Material. Holotype. NEW ZEALAND: Mid Canterbury: 1♂ ( BMNH), glued on rectangular card, “type” [red label, printed], “3695. ♂ ” [white label, handwritten], “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed], “Pudding Hill. 10.12.1912.” [white label, handwritten]; “ Sagola socia . ♂.” [white label, handwritten]. One of two syntypes of Sagola unicalis . NEW ZEALAND: Mid Canterbury: 1♂ ( BMNH), glued on rectangular card, “3826. ♂ ” [white label, handwritten], “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed], “Moa Basin 20.10.1913.” [white label, handwritten]; “ Sagola unicalis ” [white label, handwritten].

Additional Material (n = 9: 8 males; 1 female). NEW ZEALAND: Fiordland : 1♂, Barrier River, 1067m, little Red Hill , 2 II 1975, G.W. Ramsay, litter 75/50 ; 1♂, W. Olivine Ra., Tempest Spur Summit , 1463m, 30 I 1975, G.W. Ramsay, litter 75/46 ; 1♂, Middle basin, Tutoko bench, Darran mts , 1524m, 14 I 1977, J.S. Dugdale, swards 77/9; Mid Canterbury: 2♂♂ (1♂, slide-mounted), Arthurs Pass, Temple Basin , 1493m, 8 II 1982, J.S. Dugdale, sedges and mats 82/24 ; Westland: 1♂ 1♀, Mt Aspiring NP, Arawata Biv , 840m, 5 II 1989, LCNZ 89 /2, J.W. Early, R. M. Emberson, scrub litter ; 1♂, Chancellor Hut, Fox Glacier , 29 I 2005, J. Nunn, in tussock litter at 1140m ; 1♂, Alex Knob, 1280m, nr Franz Josef , 6 II 1984, J.C. Watt, turf 84/15 .

Diagnosis. This species can be distinguished from the other species of the socia species-group by the following combination of characters: larger body, length 2.5–2.8 mm; male antennomere 7 enlarged and longer than wide; eye prominent, one-third length of temple; male hind wings welldeveloped, female hind wings reduced to small pads; shape of genitalia unique to species.

Redescription. Length 2.5–2.8 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler ( Fig. 47a ). Head: As long as wide, widest across eyes ( Fig. 47k ). Ventral surface of head weakly convex. Male antennomere 1 approximately 2 times longer than wide, 2–6 subquadrate, 4–7 enlarged, 7 distinctly enlarged and longer than wide, 8–10 subquadrate, 4–11 bearing tubercles. Female antennomeres 2–10 subquadrate, 4–11 bearing tubercles. Frontal sulcus deep, reaching posterior end of eye ( Fig. 47k ). Anterior frontal fovea oval and partially covered by frontal rostrum ( Fig. 47k ), posterior frontal fovea oval ( Fig. 47k ). Eye large and prominent, approximately one-third length of temple ( Fig. 47k ). Thorax: Prosternum as long as wide, widest at midpoint. Male elytra rectangular ( Fig. 47a ), female elytra subquadrate. Male hind wings well-developed, female hind wings reduced to small pads. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe divided vertically, minor lobe broader and triangular apically ( Fig. 47f ). Phallobase of median lobe symmetrical and rounded ( Fig. 47f ). Parameres symmetrical with setae apically ( Fig. 47f ).

Type Locality. Pudding Hill , near Methven, Mid Canterbury, New Zealand .

Distribution. F i o r d l a n d, M i d C a nt e r b ur y, Westland ( Fig. 48 : black circles).

Habitat. Specimens of this species were collected by sifting forest litter at high elevations.

Comments. We could not examine one of the two syntypes of S. unicalis , but they share a unique enlarged male antennomere 7 that is longer than wide, a diagnostic species character for S. socia . One specimen was collected at the same locality as S. socia and was identified as S. unicalis by T. Broun, the author of the species, and additional specimens have been collected near the type locality. For these reasons, we have placed S. unicalis in synonymy with S. socia .

Figures

Fig. 47. Sagola species in the socia species-group. Habitus, dorsal views: a) S. socia, b) S. solodovnikovi, c) S. kahurangiensis, d) S. bullerensis, e) S. canterburyensis. Aedeagi, dorsal views: f) S. socia, g) S. solodovnikovi, h) S. kahurangiensis, i) S. bullerensis, j) S. canterburyensis. k) S. socia, dorsal surface of male head. Scale bars a–e = 1 mm, f–k = 0.1 mm.

Fig. 48. Known collection localities of New Zealand Sagola species. S. socia: black circles; S. solodovnikovi: triangles; S. kahurangiensis: black square; S. bullerensis: star; S. canterburyensis: white circle.