Calloeneis johnnie Gordon and Hanley, 2020

Gordon, Robert D., F, Guillermo González & Hanley, Guy A., 2020, South American Coccinellidae (Coleoptera), part XX: systematic revision of South American Calloeneis Grote (Cryptognathini), Insecta Mundi 2020 (766), pp. 1-25 : 12

13. Calloeneis johnnie Gordon and Hanley , new species

Description. Male holotype. Length 1.8 mm, width 1.6 mm; dorsal surface somewhat alutaceous; head strongly alutaceous, pronotum, elytron with faint trace of microsculpture. Color reddish brown; head yellow; pronotum with lateral 1/3 yellow, median 1/3 brown; elytron reddish brown medially, anterior and lateral borders dark brown ( Fig. 54 ); venter yellow except pro-, meso-, and metasternum brown; epipleuron reddish yellow; abdomen brownish yellow. Head with punctures not visible; pronotal punctures small, separated by a diameter or less; elytral punctures larger than on pronotum, separated by up to twice a diameter; prosternal, mesosternal punctures fine, barely visible; metasternal punc- tures small in median 1/3, separated by a diameter or less, punctures absent in lateral 1/3; abdominal ventrites 1, 2 with fine punctures separated by up to three times a diameter medially, ventrites 2–4 finely, densely punctured medially; ventrite 5 finely, densely punctured. Head with frons widened from vertex to clypeal apex, slightly wider than eye, clypeal apex arcuate ( Fig. 55 ); eye canthus short, about half width of eye. Prosternum with lateral carina on each side curved, extended from apex of intercoxal process to lateral margin of prosternum. Epipleuron not descending externally. Postcoxal line on basal abdominal ventrite curved, extended 2/3 distance to rear margin of ventrite. Genitalia with penis guide 3/4 as long as paramere, slender, slightly oval, tapered to abrupt apex in apical 1/4; paramere slender, weakly curved, apex rounded ( Fig. 56, 57 ); penis slender, apex attenuate, basal capsule wide, flat, inner arm straight, slender, apically truncated, outer arm wide, apically rounded ( Fig. 58 ).

Female. Similar to male except head reddish brown, pronotum entirely dark brown. Genitalia with ramus of spermathecal capsule bulbous, cornu curved, apex acute.

Variation. Length 1.7 to 1.8 mm, Size of median reddish-brown area on elytra varies from wide to almost entirely absent, rendering elytra nearly all black.

Type material. Holotype male: TRINIDAD, Brigand Hill , X.1966, C.I. E. A. 1306, C.I.B.C., 66/684, Pred. on Pseudaulacaspis pentagona, Pres by Comm Inst Ent BM 1971-1 ( BMNH) . Paratypes 5, same data as holotype ( BMNH).

Remarks. Calloeneis johnnie does not have a distinctive dorsal color pattern, but the head widened from base to the wide, apically rounded clypeus is thus far unknown elsewhere in the genus.

Figures

Figures 54–68. Calloeneis spp. 54–58) Calloeneis johnnie. 54) Habitus. 55) Frons. 56) Penis guide ventral. 57) Penis guide lateral. 58) Penis. 59–63) Calloeneis robyn. 59) Habitus. 60) Frons. 61) Penis guide ventral. 62) Penis guide lateral.63) Penis.64–68) Calloeneis francis. 64) Habitus.65) Frons.66) Penis guide ventral.67) Penis guide lateral. 68) Spermathecal capsule.