Eocnides fragilis Uéno, 1989

Deuve, Thierry, Kavanaugh, David H. & Liang, Hongbin, 2016, Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, Western Yunnan Province, China: Species of the Tribe Trechini (Coleoptera: Caraboidea), with Descriptions of Four New Genera, One New Subgenus and 19 New Species., Proceedings of the California Academy of Sciences 63 (12), pp. 341-455 : 377

Eocnides fragilis Uéno, 1989

( Figs. 15 , 45–48 )

Eocnides fragilis Uéno, 1989:14 . Holotype, a male, in NMST. Type locality: China, Sichuan, Nanping County, Jiuzhaigou, Xiajijie Hai , ca. 2600 m.

Agonotrechiotes longiantennatus Deuve 2010:17 . NEW SYNONYMY

DIAGNOSIS.— Adults of this species ( Fig. 15a ) can be distinguished from those of all other species in the region by the following combination of character states: size large (BL = 4.8 to 5.2 mm), fully-winged; body color pale, brownish-tan; head with eyes only moderate in size; frontal furrows attenuated; anterior pair of supraorbital setae not foveate; pronotum slightly transverse (ratio PW/PL= 1.48), median longitudinal impression distinctly impressed, especially basally, least so at middle; basal foveae prolonged anteriorly parallel to lateral margin in basal one-half; elytra with discal stria 2 partially effaced near base, striae 5 and 6 faintly but distinctly impressed; medi- an lobe of male aedeagus ( Fig. 15b ) slender, apex slightly recurved ventrally.

HABITAT DISTRIBUTION.— Members of this species were found by splashing gravel bars at the edges of a small, lowland stream adjacent to disturbed, agricultural areas, at elevations ranging from 1610 to 1630 m. This microhabitat is similar to that described by Uéno (1999b) for specimens collected at Munigou He, 2670 m, in Songpan County, northern Sichuan. No other trechines were collected at these sites.

GEOGRAPHICAL DISTRIBUTION WITHIN THE GAOLIGONG SHAN.— Fig. 15c . We examined a total of 2 specimens (1 male and 1 female) from the following localities: Gongshan County: Bingzhongluo Township ( Niwaluo He , just below Nu Jiang Road , 1630 m, N28.05140° / E98.59319°, 8 October 2002, D.H. Kavanaugh , P.E. Marek and D.Z. Dong collectors [1 male; CAS]; Bingzhongluo Township (Yimaluo He, just below Nu Jiang Road, 1610 m, N28.02499° / E98.62564°, 8 October 2002, D.H. Kavanaugh , P.E. Marek and H.B. Liang collectors [1 female; IOZ]. Both of these localities are in Core Area 2.

OVERALL GEOGRAPHICAL DISTRIBUTION.— Fig. 45 . This species has been recorded previously from five localities in northern Sichuan Province, in Jiuzhaigou and Songpan Counties ( Uéno 1989, 1999b) and in Luding County ( Deuve 2010). We report here the following new record for Sichuan: Wenchuan County: Qionglai Shan, Sanjiang Township (Sanjiang Nature Reserve, river at gate to reserve, 1300 m, N30.94697° / E103.31217°, 9 September 2007, D.H. Kavanaugh collector [1 female; CAS]. This record extends the range of E. fragilis in Sichuan. The discovery of E. fragilis in the northern Gaoligong Shan region, in western Yunnan Province, extends its known range an additional 260 km SW.

Figures

FIGURE 15. Eocnides fragilis Uéno; a. Dorsal habitus (CASENT1029485). b. Median lobe of aedeagus of male (CASENT1029485), left lateral aspect. c. Map of locality records (red circles) for E. fragilis in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km.

FIGURE 45. Map showing approximate overall known geographical distributions of species occurring in the Gaoligong Shan region as well as outside the region. 1 = Peripleptus imaicus Jeannel. 2 = Agonotrechus wuyipeng Deuve. 3 = Eoc- nides fragilis Uéno. 4 = Trechus indicus Putzeys. Scale line = 500 km.

FIGURE 46. Chart showing the representation of trechine species in project-designated Core Areas (see Fig. 3) in the Gaoligong Shan region.

FIGURE 47. Chart illustrating the altitudinal ranges of trechine species represented in the Gaoligong Shan region.

FIGURE. 48. Chart illustrating the co-occurence (syntopy) of trechine species in samples from the same habitats and at the same sites in the Gaoligong Shan region.