Pseudoblothrus roszkovskii (Redikorzev, 1918)

Kolesnikov, Vasiliy B. & Turbanov, Ilya S., 2018, The cave-dwelling false scorpion genus Pseudoblothrus Beier, 1931 (Arachnida: Pseudoscorpiones: Syarinidae) in the Crimean Peninsula, Zootaxa 4374 (4), pp. 524-544 : 527-540

Pseudoblothrus roszkovskii (Redikorzev, 1918)

( FIGS 3‒7 )

Ideoblothrus roszkovskii REDIkORzEV 1918: 94‒96, FIGS 3 , 4, 4A ; PLIGINSkY 1927: 172.

Pseudoblothrus roszkovskii (REDIkORzEV): BEIER 1931: 21‒22; BEIER 1932A: 136, FIG. 167; ROEWER 1937: 249; WOLF 1938: 618; ROEWER 1940: 345; VAcHON 1954: 217; BEIER 1963: 228, FIG. 234; VAcHON 1969: 392; BEIER 1970: 322, FIG. 2 ; MUcHmORE 1982: 218; INzAGHI 1983: 46; MAHNERT 1990: 167; HARVEY 1991: 428; ZARAGOzA 2010: 52, FIG. 28; GARDINI 2015: 32; KOLESNIkOV & TURbANOV 2015: 83; TURbANOV et al. 2016b: 1286; KOLESNIkOV & TURbANOV 2017: 50‒55, FIG. 1 .

Pseudoblothrus ljovuschkini KRUmPáL 1984: 642‒645, FIGS 11‒16; HARVEY 1991: 428; GARDINI 2015: 32; KOLESNIkOV & TURbANOV 2015: 82. SYNONYmIzED bY KOLESNIkOV & TURbANOV (2017).

Obisium tauricum LEbEDEV 1927: 44 ‒47, FIGS 1‒2 ; TURbANOV & KOLESNIkOV 2015: 83; TURbANOV et al. 2016b: 1286. SYNONYmIzED bY KOLESNIkOV & TURbANOV (2017).

Obisium touricum [SIc], BEIER 1932b: 277; HARVEY 1991: 671.

Type material examined. PARATуPES OF Pseudoblothrus ljovuschkini KRUMPáL, 1984, 2 MALES (ZMUM TI-1, TI-2), 1 FEMALE (ZMUM TI-3) “USSR, СRIMEA, KARAbI KARST MASSIF, EGIZ-TINAcH-2 СAVE, V.1964, LEG. V.V. ILуUcKHIN”.

Other type material. AccORDING TO REDIKORZEV (1918), SуNTуPES OF Ideoblothrus roszkovskii , 1 ADULT FEMALE AND 2 SUbADULT FEMALES ( ZISP 208), FROM ‘СRIMEA, LOWER PLATEAU OF TSHATуR-DAGH, SUUK-KObA СAVE, 7.VII.1916, V. ROSZKOVSKу’, SHOULD bE HOUSED IN THE ZOOLOGIcAL INSTITUTE OF THE RUSSIAN AcADEMу OF ScIENcES IN ST. PETERSbURG, THIS bEING ALSO cONFIRMED bу THE ZISP AccESSION NUMbER (208). HOWEVER, NO TRAcE OF THE TуPE SERIES cOULD bE FOUND IN ZISP AS OF MAу 2017.

THE TуPE MATERIAL OF Obisium tauricum , DESPITE OUR EFFORTS TO RELOcATE IT IN THE ZMUM AND ZISP cOLLEcTIONS, SEEMS TO bE LOST.

Non-type material examined. СRIMEA, Tshatyr-Dagh Karst Massif: 1 MALE (ZMUM TI-43), 1 MALE (VK), ALUSHTINSKAуA СAVE, 11.02.2015, LEG. I.S. TURbANOV; 1 MALE, 1 FEMALE (ZMUM TI-44), PASKHALNAуA СAVE, 12.02.2015, LEG. I.S. TURbANOV; 1 FEMALE, 1 TRITONуMPH (ZMUM TI-45), 1 FEMALE (VK), SUUK-KObA (= KHOLODNAуA) СAVE, 13.02.2015, LEG. I.S. TURbANOV; 1 TRITONуMPH (VK), OZIORNAуA СAVE, 13.02.2015, LEG. I.S. TURbANOV; 1 MALE (ZIPS 1432), TRIOKHETAZHNAуA СAVE, 0 8.01.2015, LEG. A.A. FROLOVA; 1 MALE (VK), EMINE-BAIR- KObA (= TRIOKHGLAZKA) СAVE, 14.02.2015, LEG. I.S. TURbANOV. Dolgorukovsky Karst Massif: 1 MALE (ZMUM TI-46), KIZIL-KObA (= KRASNAуA) СAVE, 0 5.11.2014, LEG. I.S. TURbANOV; 1 FEMALE (ZMUM TI-47), VOSTOTSHNу POTOK СAVE, 13.05.2015, LEG. I.S. TURbANOV; 1 FEMALE (ZMUM TI-48), PARTIZANSKAуA СAVE, 14.05.2015, LEG. I.S. TURbANOV. Karabi Karst Massif: 1 MALE (VK), TUAKSKAуA (= FUL-KObA) СAVE, 0 8.05.2013, LEG. I.S. TURbANOV; 1 MALE (ZMUM TI-49), TUAKSKAуA (= FUL-KObA) СAVE, 0 7.06.2014, LEG. I.S. TURbANOV; 1 MALE (IT), EGIZ-TINAKH-2 (= PASTUSHуA) СAVE, 11.06.2016, LEG. I.S. TURbANOV.

Type locality of P. roszkovskii . SUUK-KObA (= KHOLODNAуA) СAVE, TSHATуR-DAGH KARST MASSIF, СRIMEA.

Type locality of P. ljovuschkini . EGIZ-TINAKH-2 (= PASTUSHуA) СAVE, KARAbI KARST MASSIF, СRIMEA.

Type locality of O. tauricum . EMINE-BAIR-KObA (= TRIOKHGLAZKA), TSHATуR-DAGH KARST MASSIF, СRIMEA.

Diagnosis (adults). Pseudoblothrus SPEcIES DIFFERING FROM OTHER MEMbERS OF THE GENUS bу THE FOLLOWING cOMbINATION OF cHARAcTERS: WITHOUT EуES OR EуE-SPOTS; cARAPAcE WITH SIх (RARELу FIVE) SETAE IN ANTERIOR ROW AND 4‒5 IN POSTERIOR ROW; MALE WITH VENTRAL GLANDS ON STERNITE VI (3‒4 PATcHES OF GLANDULAR AREAS AND 12‒22 VENTRAL GLAND SETAE); STERNITES III AND IV WITH 3‒5 AND 3‒6 STIGMATAL SETAE ON EAcH SIDE, RESPEcTIVELу; PEDIPALP GRANULATE; cHELA LONG AND NARROW, HAND WITH SUbPARALLEL EDGES; ist PLAcED cLOSE TO est (DISTANcE bETWEEN THEM NOT MORE THAN 0.25 MM IN DORSAL ASPEcT). LENGTH OF PEDIPALPAL FEMUR, 1.70‒ 2.10 MM (6.64‒8.07 TIMES AS LONG AS bROAD), OF PATELLA, 1.50 ‒1.88 MM (4.54‒6.0×), OF cHELA WITH PEDIcEL, 2.60‒3.24 MM (6.04‒6.63×); FIхED AND MOVAbLE cHELAL FINGERS WITH 120‒126 AND 110‒118 TEETH, RESPEcTIVELу.

Description of adults (male and female). AMONG THE LARGEST REPRESENTATIVES OF THE GENUS Pseudoblothrus ( TAbLES 3‒4), WITH cHARAcTERISTIc TROGLOMORPHIc FEATURES ( FIG. 3 ). СARAPAcE, PEDIPALPS AND TERGITES I‒II REDDISH bROWN, TERGITES II‒XI PALE bROWN. OPISTHOSOMA AND LEGS PALE.

СARAPAcE ( FIGS. 4A‒E ). 1.25‒1.51 LONGER THAN WIDE, WITH TWO TRANSVERSE FURROWS, WITHOUT EуES OR EуE SPOTS. ANTERIOR MARGIN WITHOUT EPISTOME, SLIGHTLу cURVED FORWARD, bASE OF cHELIcERAL TEcTUM (λ) bENT AT AN ObTUSE ANGLE ( FIG. 4F ). TEGUMENT cOVERED WITH A WEAK MESHWORK OF ELONGATED REcTANGULAR cELLS. SETAE THIN, ARRANGED IN FIVE MAIN ROWS (SOME SPEcIMENS WITH ADDITIONAL ROWS), cHAETOTAху 5‒6:4(0‒2):4:3‒5(0‒4):4‒5 (TOTAL 21‒28), NUMbER AND LOcATION OF SETAE VARIAbLE, OFTEN WITH A MARKED ASуMMETRу. СHAETOTAху OF cARAPAcE IN SPEcIMENS OF EAcH SEх FROM DIFFERENT KARST MASSIFS AS PRESENTED IN TAbLE 5 . THREE PAIRS OF LуRIFISSURES: TWO PAIRS bETWEEN ROWS 1 AND 2 AND ONE PAIR bETWEEN ROWS 4 AND 5, RESPEcTIVELу. NUMEROUS MINUTE PORES ALL OVER cARAPAcE.

СOхAE ( FIG. 4R ). MANDUcATORу PROcESS TRIANGULAR AND AcUMINATE, WITH TWO SETAE, EAcH 0.2‒0.3 MM LONG. PEDIPALPAL cOхA cOVERED WITH A FINE MESH ScULPTURING. СHAETOTAху AS IN TAbLE 5.

OPISTHOSOMA ( FIGS 4O‒Q ). PLEURAL MEMbRANE LONGITUDINALLу STRIATE ( FIG. 4J ). TERGAL AND STERNAL cHAETOTAху IN SPEcIMENS FROM DIFFERENT KARSTS AND IN DIFFERENT SEхES AS PRESENTED IN TAbLE 5. MALE GENITAL OPENING AS IN FIGS. 4K‒ M . MALE STERNITES IV‒V WITH A GROUP OF SMALL MEDIAN SETAE ( FIGS. 4I, 4P ). VENTRAL GLAND OF MALE WITH 3‒4 GLANDULAR AREAS ( FIG. 4N ). FEMALE STERNITES IV‒V WITHOUT MEDIAN MIcROSETAE, bUT 0‒2 POSTERIOR SETAE MAу bE SHORTER THAN THE REST ( FIG. 4Q ). STIGMATA OF STERNITES III AND IV AS IN FIG. 4S . ONE FEMALE WITH 4 SETAE ON TERGITE XII (TERATOLOGу).

GENITALIA. MALE GENITALIA AS IN FIG. 4G , GENITAL cHAMbER WITH TWO UNMODIFIED SETAE; MEDIAN GENITAL SAc WITH TWO bRANcHES. FEMALE GENITALIA AS IN FIG. 5H .

СHELIcERA ( FIG. 5I ). HAND WITH FIVE SETAE. GALEAL SETA (gs) 0.69‒0.73 FROM bASE OF MOVAbLE FINGER. GALEA AbSENT IN bOTH SEхES. FIхED FINGER WITH 13‒17 LARGE AND 3‒7 SMALL TEETH. MOVAbLE FINGER WITH 8‒12 PRIMARу TEETH, OF WHIcH 2‒3 LARGER THAN OTHERS, AND 2‒4 SMALL TEETH. LуRIFISSURES LOcATED AT bASE OF SETAE is. RALLUM WITH 7‒8 bLADES, EAcH 0.16‒0.18 MM LONG AND FINELу SERRATE ON ANTERIOR EDGE ( FIG. 5J ). SERRULA EхTERIOR WITH 25 bLADES ( FIG. 5K ), SERRULA INTERIOR WITH 16 ( FIG. 5L ). HAND AND bASE OF FINGERS RETIcULATE-GRANULATE. MEASUREMENTS OF cHELIcERA AS IN TAbLE 3.

PEDIPALP ( FIGS. 5A ‒С, 6). TROcHANTER WITH cOARSE, DENTIFORM GRANULATION; FEMUR, PATELLA, HAND AND bASE OF FINGERS WITH UNIFORM, ROUNDED GRANULATION. BASE OF cHELAL PEDIcEL MORE DENSELу AND cOARSELу GRANULATED. TROcHANTER USUALLу WITH 1‒2 SHORT TUbERcLES EcTALLу, IN SOME SPEcIMENS THESE cAN bE AbSENT. FEMUR LONG, GRADUALLу bROADENING DISTALLу. PATELLA cLAVATE, SLIGHTLу EхPANDED ON MESAL SIDE AT 0.3 OF LENGTH FROM bASE, WHERE FOUR LуRIFISSURES OF DIFFERENT SIZES ARE LOcATED, TWO OF THEM cLOSE TOGETHER. СHELA LONG AND NARROW, HAND WITH SUbPARALLEL EDGES IN DORSAL AND LATERAL VIEWS. FINGERS SLIGHTLу cURVED WHEN VIEWED FROM AbOVE. ALTOGETHER, 12 TRIcHObOTHRIA; FIхED FINGER WITH 4 IN DISTAL PART OF FIхED FINGER (et, it, ist AND est), 2 IN bASAL PART OF FIхED FINGER (esb AND isb), AND 2 ON DISTAL PART OF HAND (ib AND eb). TRIcHObOTHRIOTAху AS IN FIGS. 6A‒F . DISTANcES bETWEEN bOTHRIDIA AS IN TAbLE 6 . HAIRS OF TRIcHObOTHRIA LONG, SIMPLE AND FILIFORM, TRIcHObOTHRIUM t NOT LANcEOLATE. DORSAL SURFAcE OF cHELA, bETWEEN FINGER AND ib, WITH 2‒3 IRREGULAR ROWS OF MIcROSETAE (cHEMOSENSORу SETAE, sc), EcTAL IN POSITION. MIcROSETAE ALSO ObSERVED OVER MOST OF bOTH FINGERS, ADJAcENT TO DENTAL MARGIN. СOUPLED SENSILLA (pc) PRESENT ON MOVAbLE FINGER bETWEEN st AND sb, NEAR DENTAL MARGIN. LуRIFISSURES ARRANGED AT bASE OF HAND ON DORSAL SIDE, bETWEEN LEVELS OF b AND esb, bETWEEN esb AND isb, bETWEEN b AND sb (1‒2 LуRIFISSURES), AND bETWEEN LEVELS OF est AND it (ON DORSAL SIDE). FIхED FINGER WITH 120‒126 SMALL, ALMOST POINTED, cONTIGUOUS TEETH. TEETH OF FIхED FINGER REAcHING TO LEVEL OF isb. MOVAbLE FINGER WITH SIMILAR TEETH, 110‒118 IN NUMbER. TEETH OF MOVAbLE FINGER NOT REAcHING AS FAR bASALLу AS THOSE OF FIхED FINGER (DIFFERENcE EqUIVALENT TO AbOUT 4‒5 TEETH). FIхED FINGER WITH A WELL DEVELOPED VENEDENS AND VENOM GLAND . Lamina defensor (ld) APPRESSED TO VENEDENS. VENOM DUcT SHORT, nodus ramosus WELL DISTAD OF et ( FIG. 5D ). MOVAbLE FINGER WITHOUT VENOM APPARATUS, APODENS SHORT, WITH A SMALL lamina defensor ( FIG. 5E ).

LEGS I AND IV ( FIGS. 5F, G ). SMOOTH, EхcEPT FOR TROcHANTER AND FEMUR, WHIcH ARE cOVERED WITH A FINE MESH ScULPTURING. FEMUR OF LEG I 1.48‒2.0× LONGER THAN PATELLA. FEMUR OF LEG IV 0.45‒0.56× AS LONG AS PATELLA; FEMUR+PATELLA 1.0‒1.23× LONGER THAN TIbIA. JOINT bETWEEN FEMUR AND PATELLA OF LEG IV PERPENDIcULAR. TELOTARSI NOT EхPANDED, GRADUALLу TAPERING DISTALLу. SUbTERMINAL TARSAL SETAE OF ALL LEGS SERRATE DISTALLу, EAcH WITH 2‒3 bARbS ( FIG. 5H ). СLAWS SMOOTH, THIN, ARcUATE; AROLIA UNDIVIDED AND DISTINcTLу SHORTER THAN cLAWS ( FIG. 5M ).

MEASUREMENTS AND RATIOS OF ADULTS AS IN TAbLES 3‒4.

Description of tritonymph. СARAPAcE, PEDIPALPS, PEDIPALPAL cOхAE AND cHELIcERAE PALE REDDISH bROWN, OTHER PARTS PALE уELLOW.

СARAPAcE ( FIG. 7O ). 1.28‒1.31× LONGER THAN WIDE, WITHOUT EуES OR EуE SPOTS, WITH TWO TRANSVERSE FURROWS. EPISTOME AbSENT; bASE OF cHELIcERAL TEcTUM bENT AT AN ObTUSE ANGLE MEDIALLу ( FIG. 7P ). СHAETOTAху OF cARAPAcE 6:4:4:3‒4:4 (21‒22). THREE PAIRS OF LуRIFISSURES: TWO PAIRS bETWEEN ROWS 1 AND 2, ONE PAIR bETWEEN ROWS 4 AND 5. СARAPAcE cOVERED WITH A FINE MESH-LIKE ScULPTURING. NUMEROUS MINUTE PORES OVER ENTIRE cARAPAcE.

СOхAE ( FIG. 7N ). MANDUcATORу PROcESS TRIANGULAR, AcUMINATE, WITH TWO SETAE. PEDIPALPAL cOхA cOVERED WITH A FINE MESH ScULPTURING. SURFAcE OF cOхAE I‒IV SMOOTH. СHAETOTAху FOR DIFFERENT POPULATIONS GIVEN IN TAbLE 5.

OPISTHOSOMA. PLEURAL MEMbRANE LONGITUDINALLу STRIATE ( FIGS. 7L‒M ). TERGAL AND STERNAL cHAETOTAху FOR DIFFERENT POPULATIONS GIVEN IN TAbLE 5. STERNITE VI WITH 1‒2 MEDIAN MIcROSETAE.

СHELIcERA ( FIG. 7F ). HAND WITH FIVE SETAE. GALEAL SETA (gs) 0.66 FROM bASE OF MOVAbLE FINGER. GALEA AbSENT. FIхED FINGER WITH 14‒16 TEETH. MOVAbLE FINGER WITH 10‒14 TEETH. MIcROLуRIFISSURES LOcATED AT bASE OF SETA is. RALLUM WITH SIх bLADES, EAcH WITH A FINELу SERRATED ANTERIOR EDGE. SERRULA EхTERIOR WITH 24 bLADES ( FIG. 7G ), SERRULA INTERIOR WITH 16 bLADES. POSTERIOR EDGE OF bLADES OF SERRULA INTERIOR SERRATE ( FIG. 7H ). HAND AND bASE OF FINGERS RETIcULATE-GRANULATE.

PEDIPALP ( FIGS. 7A ‒С). TROcHANTER cOARSELу GRANULATED; FEMUR, PATELLA, HAND AND bASES OF FINGERS WITH UNIFORM, ROUNDED GRANULATION. PEDIcEL OF cHELA cOARSELу GRANULATED. TROcHANTER WITHOUT TUbERcLES ON EcTAL FAcE. TROcHANTER WITH ONE LуRIFISSURE AT DISTAL END. FEMUR LONG, GRADUALLу bROADENING DISTALLу. PATELLA cLAVATE, SLIGHTLу EхPANDED ON MESAL SIDE 0.27 OF LENGTH FROM ITS bASE, AT WHIcH POINT THERE ARE FOUR LуRIFISSURES OF DIFFERENT SIZES, TWO OF THEM cLOSE TOGETHER. СHELA LONG AND NARROW. HAND WITH MARGINS SUbPARALLEL IN DORSAL AND LATERAL VIEWS. FINGERS SLIGHTLу cURVED WHEN VIEWED FROM AbOVE. TEN TRIcHObOTHRIA: FIхED FINGER WITH 4 (et, it, ist AND est) IN DISTAL PART, 1 (esb) IN bASAL PART OF FIхED FINGER, 2 (ib AND eb) IN DISTAL PART OF HAND, TRIcHObOTHRIUM isb AbSENT; MOVAbLE FINGER WITH 3 TRIcHObOTHRIA, sb AbSENT. TRIcHObOTHRIOTAху AS IN FIGS. 7A‒B . DISTANcES bETWEEN bOTHRIDIA AS IN TAbLE 6 . HAIRS OF TRIcHObOTHRIA LONG, FILIFORM, t NOT LANcEOLATE. SETAE OF cHELA THIN. DORSAL SURFAcE OF cHELA WITH 2‒3 IRREGULAR ROWS OF MIcROSETAE (cHEMOSENSORу SETAE, sc), EcTAL IN POSITION bETWEEN ib AND bASE OF FIхED FINGER. MIcROSETAE ALSO ObSERVED ALMOST ALL OVER bOTH FINGERS, ADJAcENT TO DENTAL MARGIN. СOUPLED SENSILLA (pc) PRESENT ON MOVAbLE FINGER bETWEEN sb AND st NEAR DENTAL MARGIN. LуRIFISSURES ARRANGED AT bASE OF HAND ON DORSAL SIDE, bETWEEN b AND esb, PROхIMAL TO esb, bETWEEN b AND st (TWO MIcROLуRIFISSURES), AND bETWEEN est AND it ON DORSAL SIDE. FIхED FINGER WITH 92‒95 SMALL, cONTIGUOUS TEETH. TEETH OF FIхED FINGER REAcHING TO LEVEL OF esb. MOVAbLE FINGER WITH 88‒90 SMALL, cONTIGUOUS TEETH. TEETH OF MOVAbLE FINGER NOT EхTENDING AS FAR bASALLу AS THOSE OF FIхED FINGER (DIFFERENcE EqUIVALENT TO AbOUT 4‒5 TEETH). A ROW OF FOUR VERу SMALL DENTIcLES AT bASE OF FIхED FINGER , THESE WITHOUT DENTAL cANALS. FIхED FINGER WITH WELL DEVELOPED VENEDENS AND VENOM GLAND, VENOM DUcT SHORT, nodus ramosus cLEARLу DISTAL TO et ( FIG. 7D ). Lamina defensor (ld) TIGHTLу APPRESSED TO VENEDENS. MOVAbLE FINGER WITHOUT VENOM APPARATUS, WITH A SHORT, bLUNT APODENS AND A SMALL lamina defensor ( FIG. 7E ).

LEGS I AND IV ( FIGS. 7I ‒J ). SMOOTH, EхcEPT TROcHANTER AND FEMUR, WHIcH ARE cOVERED WITH A FINE MESH-LIKE ScULPTURING. FEMUR OF LEG I 1.53‒1.64× LONGER THAN PATELLA. FEMUR OF LEG IV HALF AS LONG AS PATELLA; FEMUR+PATELLA 1.08‒1.18× LONGER THAN TIbIA. JOINT bETWEEN FEMUR AND PATELLA OF LEG IV PERPENDIcULAR. TELOTARSI OF ALL LEGS SLIGHTLу DILATED AT bASE, THEREAFTER GRADUALLу TAPERING DISTAD. SUbTERMINAL TARSAL SETAE OF ALL LEGS SERRATE DISTALLу, EAcH WITH 2‒ 3 bARbS. СLAWS SMOOTH, THIN, ARcUATE; AROLIA UNDIVIDED AND DISTINcTLу SHORTER THAN cLAWS ( FIG. 7K ).

Chitrella cala and Syarinus sp.

……continued on the next page Tshatyr-Dagh Dolgorukovsky Karabi Overall

tritonymph female male male female male male female

Carapace 6:4:4:354:4 556:4:4:354(2):4 6:4:4:455:4 6:4:4:4(4):4 6:4:4:4(152):5 556:4(05 556:4(052):4:455(05 556:4:4:354(152):45

2):4:4(052):455 4):455 5

number of setae in 21522 20524 22523 26 24525 21528 21528 21525 carapace

Pedipalpal coxa (including 2:356:355 2:455:556 2:355:455 2:3:5 2:354:556 2:4:455 2:355:455 2:355:556 manducatory process)

Pedal coxa I 657 658 758 759 7510 859 759 6510 Pedal coxa II 657 859 759 859 859 9510 7510 859 Pedal coxa III 355 354 455 4 455 455 455 355 Pedal coxa IV 558 9512 9511 9510 9510 9511 9511 9512 Tergite I 4 455 4 5 456 4 455 456 Tergite II 6 556 6 6 6 6 6 556 Tergite III 657 7 658 6 658 7 658 758 Tergite IV 859 758 758 7 8 758 758 758 Tergite V 9 10 9510 9 9510 859 8510 9510 Tergite VI 9 9510 9510 9 9511 9 9510 9511 Tergite VII 10511 9511 9510 10 9510 9510 9510 9511 Tergite VIII 9510 9512 9511 10 9 9 9511 9512 Tergite IX 10511 10 9511 9 9 9 9511 9510 Tergite X 758 759 758 9 758 7 759 759 Tergite XI 4 455 456 6 556 556 456 456 Sternite II 254 12519 24528 29 13515 27 24529 12519 Anterior margin of genital 5 5 9511 7 5 8511 7511 5 opening on sternite II

Posterior margin of genital 5 5 658 6 5 456 458 5 opening on sternite III

Sternite III 17520 27531 37549 37 30531 44547 37549 27531 Stigmata III (per side) 253 455 355 4 455 455 355 455 Sternite IV 17518 19522 28533 26 19520 22524 22533 19522 Stigmata IV (per side) 354 355 456 4 455 4 456 355 Sternite V 15 14517 30532 30 14 25529 25532 14517 Sternite VI 16 (152) 16520 33542 29 16 30 29542 16520 Glandular area/posterior row 5 5 17522/16520 15/14 5 12514/16518 12522/14520 5 sternite VI (male)

Sternite VII 15 17518 16521 17 16 17518 16521 16518 Sternite VIII 15517 16520 16519 15 16 15517 15519 16520 Sternite IX 14515 14518 14518 15 14 14 14518 14518 Sternite X 12515 14 14516 13 11512 13514 13516 11514 Sternite XI 6 6 456 4 455 455 456 456

cone 2+2 254+2 2+2 2+2 2+2 2+2 2+2 2+2 MEASUREMENTS AND RATIOS OF TRITONуMPHS AS IN TAbLES 3‒4.

PROTONуMPH AND DEUTONуMPH UNKNOWN.

Remarks. KRUMPáL (1984), WHEN DEScRIbING P. ljovuschkini , POINTED OUT THE FOLLOWING TWO cHARAcTERS AS DIAGNOSTIc: (1) LENGTH/WIDTH RATIO OF PEDIPALP FEMUR, 7.03‒7.51 IN P. ljovuschkini , VERSUS 8.0‒8.05 IN P. roszkovskii ; LENGTH/WIDTH RATIO OF PATELLA, 4.74‒5.45 IN P. ljovuschkini , VERSUS 6.2‒6.27 IN P. roszkovskii ; (2) TERGAL cHAETOTAху 4‒ 4‒4‒4‒6‒8... IN P. ljovuschkini , VERSUS 4‒5‒6‒8‒10 ‒10... IN P. roszkovskii . GARDINI (2015), IN A KEу TO ADULTS OF ALL Pseudoblothrus SPEcIES, ALSO DREW ATTENTION TO SIZE DIFFERENcES AND THE cHELAL LENGTH/bREADTH RATIO: P. roszkovskii WAS LARGER AND MORE ATTENUATE (LENGTH OF FEMUR 2.08 MM, LENGTH OF PATELLA 1.81‒1.88 MM, LENGTH OF HAND 1.41‒1.42 (3.2× bREADTH)) THAN P. ljovuschkini (LENGTH OF FEMUR 1.77‒1.95 MM, LENGTH OF PATELLA 1.52‒1.74 MM, LENGTH OF HAND, 1.07‒1.25 MM (2.3‒2.9× bREADTH)).

THE SPEcIMENS cOLLEcTED bу US FROM TSHATуR-DAGH AND KARAbI KARST MASSIFS DO NOT ALWAуS cORRESPOND TO WITH THE INDIcATORS LENGTH/WIDTH RATIO OF PEDIPALP FEMUR AND PATELLA FROM PUbLIcATIONS KRUMPáL (1984) AND GARDINI (2015). THE MINIMUM VALUE FOR THE LENGTH/WIDTH RATIO OF THE FEMUR OF THE NEW SAMPLES FROM THE TSHATуR-DAGH KARST MASSIF (7.32×) FALLS WITHIN THE LIMITS INDIcATED bу KRUMPáL (1984) AND GARDINI (2015) FOR THE KARAbI KARST MASSIF (7.03‒7.51×). THE MINIMUM VALUE FOR THE LENGTH/WIDTH RATIO OF THE PATELLA OF THE NEW SAMPLES FROM THE TSHATуR- DAGH KARST MASSIF (5.38×) FALLS WITHIN THE LIMITS INDIcATED bу KRUMPáL (1984) AND GARDINI (2015) FOR THE KARAbI KARST MASSIF (4.74‒5.45×). WE ALSO FOUND OVERLAPPING VALUES OF THE LENGTH/WIDTH RATIO OF THE PEDIPALP PATELLA bETWEEN THE NEW SAMPLES: THE MINIMUM VALUE FOR SAMPLES FROM TSHATуR-DAGH KARST MASSIFS (5.38×) LIES WITHIN THE LIMITS ObTAINED bу US FOR THE KARAbI KARST MASSIF (5.03‒5.50×). THUS, IN bOTH OF THE INVESTIGATED KARST MASSIFS (TSHATуR-DAGH AND KARAbI) THERE ARE INDIVIDUALS POSSESSING THE FEATURES OF bOTH SPEcIES ( P. roszkovskii AND P. ljovuschkini ). OVERLAPPING VALUES OF MEASUREMENTS OF THE NEW MATERIAL IS ALSO ObSERVED FOR OTHER bODу PARTS ( TAbLES 3‒4). ALSO, WE ObSERVED OVERLAP OF THE MEASUREMENTS FOR OTHER PARTS OF THE bODу WHEN cOMPARING OUR SPEcIMENS FROM THE DOLGORUKOVSKу KARST MASSIF WITH OUR SPEcIMENS FROM THE TSHATуR-DAGH AND KARAbI KARST MASSIFS, AND WITH THE DATA IN KRUMPáL (1984) AND GARDINI (2015). THUS, THE VALUES ARE INVALID FOR SEPARATING REPRESENTATIVES OF THE GENUS Pseudoblothrus INTO TWO DIFFERENT SPEcIES, LIVING IN TSHATуR-DAGH AND KARAbI KARST MASSIFS.

KRUMPáL (1984) ALSO NOTED A SLIGHT DIFFERENcE IN THE RALLUM, WITH THAT OF P. roszkovskii HAVING SEVEN bLADES, AS OPPOSED TO EIGHT IN P. ljovuschkini . THE SPEcIMENS cOLLEcTED bу US FROM THE KARAbI KARST MASSIF HAVE SEVEN bLADES. WE FOUND EIGHT bLADES ONLу FOR ONE SPEcIMEN FROM THE TSHATуR-DAGH KARST MASSIF (OTHERS FROM THIS MASSIF HAD SEVEN bLADES). WE ASSUME THAT THE DIFFERENcE IN THE NUMbER OF bLADES REFLEcTS INTRASPEcIFIc VARIAbILITу. VARIATION IN THE NUMbER OF bLADES HAS PREVIOUSLу bEEN NOTED FOR OTHER SPEcIES OF THE GENUS (GARDINI 2015).

VARIAbILITу IS ALSO ObSERVED IN OTHER cHARAcTERS. IN THE cASE OF THE cARAPAcE cHAETOTAху, THIS IS EхPRESSED IN THE APPEARANcE OR DISAPPEARANcE OF ROWS OF SETAE, ESPEcIALLу THE INTERMEDIATE ROWS IN THE cENTRAL PART OF THE cARAPAcE. VARIATIONS ARE ALSO SEEN IN THE NUMbER OF SETAE IN THE ANTERIOR AND POSTERIOR ROWS, bUT THE DIFFERENcE THERE NEVER AMOUNTS TO MORE THAN ONE SETA, AS PREVIOUSLу NOTED bу KRUMPáL (1984). СHAETOTAcTIc VARIATIONS cAN ALSO cONcERN OTHER PARTS OF THE bODу : THE GENITAL OPENING OF THE MALE AND FEMALE, THE TERGITES AND STERNITES, THE cOхAL AREA, AND THE cHELA (INcLUDING MIcROSETAE), SHOWING SIMILAR DEGREES OF VARIATION bETWEEN DIFFERENT KARSTS AND bETWEEN cAVES OF A SINGLE KARST AREA. IN ALL cASES THE VARIAbILITу IS LIMITED TO MINOR DIFFERENcES IN THE NUMbER OF SETAE AND THEIR LOcATIONS RELATIVE TO EAcH OTHER. THE SAME IS THE cASE FOR THE LуRIFISSURES.

A SLIGHT SHIFT, WITHIN 0.10‒0.27 MM, IN THE LOcATION OF TRIcHObOTHRIA ON THE cHELA (FOR EхAMPLE, isb RELATIVE TO esb, AND OF ist RELATIVE TO est) cAN ALSO bE NOTED. THESE VARIATIONS ARE PARTLу ATTRIbUTAbLE TO DIFFERENcES IN THE SIZE OF THE cHELA (THE LENGTH OF THE cHELA+ VARIES WITHIN 2.60‒3.24). THE GENERAL ARRANGEMENT OF TRIcHObOTHRIA ON THE cHELA REMAINS WELL DEFINED, SO THE VARIATIONS cAN bE ATTRIbUTED TO INTRASPEcIFIc VARIAbILITу.

Figures

FIGURE 1. MAP OF cOLLEcTION SITES (cAVES) FOR Pseudoblothrus roszkovskii IN THE СRImEAN MOUNTAINS: 1—SUUk-KObA; 2— EmINE-BAIR-KObA; 3—TRIOkHETAzHNAYA; 4—OzIORNAYA; 5—PASkHALNAYA; 6—ALUSHTINSkAYA; 7—KIzIL-KObA; 8— PARTIzANSkAYA;9—VOSTOTSHNY POTOk;10—EGIz-TINAkH-2;11—TUAkSkAYA.

FIGURE 2. PHYLOGENY OF Pseudoblothrus roszkovskii POPULATIONS, bASED ON ANALYSIS OF mITOcHONDRIAL СOI GENE SEqUENcES USING THE NEIGHbOUR JOINING (NJ) mETHOD;bOOTSTRAP SUPPORT FOR bRANcH NODES SHOWN FOR 1000 REPLIcATIONS.

FIGURE 3. Pseudoblothrus roszkovskii, LIVE mALE FROm TUAkSkAYA СAVE (KARAbI KARST MASSIF, СRImEA).AFTER KOLESNIkOV & TURbANOV (2017), WITH PERmISSION.

FIGURE 4. Pseudoblothrus roszkovskii, FROm:SUUk-KObA СAVE, TSHATYR-DAGH KARST MASSIF (A‒B, F‒J, L, N‒S), VOSTOTSHNY POTOk СAVE (С), PARTIzANSkAYA СAVE (D), KIzIL-KObA СAVE, DOLGORUkOVSkY KARST MASSIF (K) AND EGIz-TINAkH-2 СAVE, KARAbI KARST MASSIF (E, M). A, MALE cARAPAcE. B, FEmALE cARAPAcE. С, FEmALE cARAPAcE. D, FEmALE cARAPAcE. E, MALE cARAPAcE. F, ANTERIOR PART OF mALE cARAPAcE.G,MALE GENITALIA,VENTRAL VIEW.H, FEmALEGENITALIA,VENTRAL VIEW.I, DIScALSETAE ON STERNITES IV AND V OF mALE.J, PLEURAL mEmbRANE OF FEmALE.K, MALE GENITAL OPENING.L, MALE GENITAL OPENING.M, MALE GENITAL OPENING.N, VENTRAL GLAND OF mALE STERNITE VI.O, TERGITES I‒XII, DORSAL VIEW,FEmALE.P,STERNITES II‒XII, VENTRAL VIEW,mALE.Q, STERNITES II‒ XII, VENTRAL VIEW,FEmALE.R, СOxAE OF PEDIPALP AND LEGS I‒IV, FEmALE.S, LEFT STIGmATA OF STERNITES III AND IV, mALE.

FIGURE 5. Pseudoblothrus roszkovskii, mALES (UNLESS OTHERWISE NOTED), FROm SUUk-KObA СAVE (TSHATYR-DAGH KARST MASSIF) (A, D‒M), VOSTOTSHNY POTOk СAVE (DOLGORUkOVSkY KARST MASSIF) (B) AND EGIz-TINAkH-2 СAVE (KARAbI KARST MASSIF) (С). A, RIGHT PEDIPALP mINUS cHELA.B, RIGHT PEDIPALP mINUS cHELA (FEmALE). С, RIGHT PEDIPALP mINUS cHELA.D, APEx OF FIxED cHELAL FINGER, EcTAL VIEW. E, APEx OF mOVAbLE cHELAL FINGER, EcTAL VIEW. F, RIGHT LEG IV. G, RIGHT LEG I. H, SUbDISTAL SETA OF LEG IV. I, RIGHT cHELIcERA.J, RALLUmOF RIGHT cHELIcERA,mESAL VIEW.K, SERRULA ExTERIOROF RIGHT cHELIcERA,EcTAL VIEW.L, SERRULA INTERIOR OF RIGHT cHELIcERA,mESAL VIEW.M, DISTAL PORTION OF TARSUS AND APOTELE OF RIGHT LEG IV.

FIGURE 6. Pseudoblothrus roszkovskii, LEFT cHELA. A, MALE, SUUk-KObA СAVE (TSHATYR-DAGH KARST MASSIF), EcTAL. B, E: FEmALE, VOSTOTSHNY POTOk СAVE (DOLGORUkOVSkY KARST MASSIF), EcTAL. С, MALE, mALE, EGIz-TINAkH-2 СAVE (KARAbI KARST MASSIF), EcTAL. D, MALE, SUUk-KObA СAVE (TSHATYR-DAGH KARST MASSIF), DORSAL. E, FEmALE, VOSTOTSHNY POTOk СAVE (DOLGORUkOVSkY KARST MASSIF),DORSAL.F, MALE,EGIz-TINAkH-2 СAVE (KARAbI KARST MASSIF),DORSAL.

FIGURE7. Pseudoblothrus roszkovskii, TRITONYmPH,OzIORNAYA СAVE (TSHATYR-DAGH KARST MASSIF).A, RIGHT cHELA,EcTAL VIEW. B,RIGHT cHELA, DORSAL VIEW.С, LEFT PEDIPALP,mINUS cHELA.D, APEx OF FIxED cHELAL FINGER,EcTAL VIEW.E, TIP OF mOVAbLE cHELAL FINGER, EcTAL VIEW. F, RIGHT cHELIcERA. G, SERRULA ExTERIOR OF RIGHT cHELIcERA, EcTAL VIEW. H, SERRULA INTERIOR OF RIGHT cHELIcERA, mESAL VIEW.I, RIGHT LEG I. J, RIGHT LEG IV.K, DISTAL PORTION OF TARSUS AND APOTELE OF RIGHT LEG IV.L, STERNITES II‒XII.M, TERGITES I‒XII.N, СOxAE OF PEDIPALP AND LEGS I‒IV. O, СARAPAcE.P, ANTERIOR PART OF cARAPAcE.