Campostichomma alawala, Polotow & Griswold, 2017

Polotow, Daniele & Griswold, Charles, 2017, Cleaning old cabinets: revealing the taxonomy of Sri Lankan wolf spiders (Araneae, Udubidae and Zoropsidae), Zootaxa 4362 (1), pp. 51-74 : 56-59

Campostichomma alawala sp. nov.

Figures 1 , 5 , 8A–B , 14

Type material. Female holotype from Sri Lanka, Alawala, 42 km (26 miles) NE Colombo, 6.III.1962, CASENT9067599, deposited in Lund University. Two female paratypes with the same data as holotype. Etymology. The species epithet is a noun in apposition from the type locality.

Diagnosis. Females of C. alawala can be distinguished by the shape of the epigynal structures, as the presence of a broad and slightly sclerotized epigynal plate, with an excavated median lobe plate and curved copulatory openings; epigynal teeth small, lateral, at outer margins of lateral lobes ( Figs 5D , 8A ); and by the copulatory ducts with one ascending spiral ( Figs 5E, F , 8B ).

DesCription. Female (holotype): Total length 4.30. Markings as in Fig. 5A–C . Carapace 2.45 long, 1.80 wide, 0.80 high; clypeus 0.10 high. Eye diameters: AME 0.09, ALE 0.16, PME 0.10, PLE 0.17. Chelicerae 0.96 long; sternum 1.20 long, 0.90 wide; labium 0.46 long; palpal coxae 0.85 long. Spination as in genus description. Leg measurements (Femur + Patella + Tibia + Metatarsus + Tarsus = [Total]): I: 1.65 + 0.88 + 1.40 + 1.30 + 0.90 = 6.13; II: 1.60 + 0.80 + 1.30 + 1.30 + 0.80 = 5.80; III: 1.50 + 0.70 + 1.00 + 1.40 + 0.85 = 5.45; IV: 1.95 + 0.75 + 1.60 + 2.45 + 1.30 = 8.05. Leg formula 4123. Epigynum: median lobe large and excavated; lateral lobes with small, lateral teeth at outer margins of lateral lobes ( Figs 5D , 8A ). Vulva with elongated and sinuous copulatory ducts, which have one ascending spiral; spermathecae large, with projections and small fertilization ducts ( Figs 5E, F , 8B ).

Variation. Female (N= 3): Total length 4.30–4.70; carapace length 2.40–2.45; femur I 1.50–1.65.

Male. Unknown.

Material examined. Only the types.

Natural history. Collection data for the specimens as described in Brinck et al. (1971: 24) refer to a low elevation site, a “hill top” at 200 m that was “covered by jungle [and] much bamboo.” C. alawala differs from other Campostichomma by occurring at low elevation: others occur in highlands above 1000 m (Brinck et al. 1971).

Distribution. Known only from the type locality at Alawala in Sri Lanka ( Fig. 14 ).

Figures

FIGURE 1. Campostichomma alawala sp. nov. (CASENT9067599), FEmAlE lEFt lEg I: (A) clAws; (B) tArsus (ArrOw pOInts tO tArsAl OrgAn); (C) mEtAtArsus, sEtAE And FEAthEry scAlEs; (D) tArsus-mEtAtArsus jOInt, shOwIng mEtAtArsAl stOppEr; (E) tArsAl OrgAn;(F) trIchObOthrIAl bAsE,tArsus.

FIGURE5. Campostichomma alawalasp. nov., FEmAlE.(A) hAbItus,dOrsAl VIEw;(B) VEntrAl VIEw; (C)AntErIOr VIEw OF OculAr ArEA;(D–F) EpIgynum; (D)VEntrAl VIEw;(E) dOrsAl VIEw; (F) lAtErAl VIEw.

FIGURE8. Campostichomma spp., FEmAlE gEnItAlIA.(A–B) C. alawala sp. nov., EpIgynum;(A) VEntrAl VIEw;(B) dOrsAlVIEw; (C–D) C. harasbedda sp. nov., EpIgynum; (C) VEntrAl VIEw; (D) dOrsAl VIEw;(E–F) C. mudduk sp. nov., EpIgynum; (E)VEntrAl VIEw; (F) dOrsAl VIEw. AbbrEVIAtIOns: CD, cOpulAtOry ducts; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; LP, lAtErAl prOcEss OF EpIgynum; ML, mEdIAn lObEOF EpIgynum;S, spErmAthEcAE.

FIGURE14. (A) PArtIAl wOrld mAp (ExcEptNEArctIc AndNEOtrOpIcAl rEAlms),shOwIng thE lOcAtIOnOF thE IslAnd cOuntry OF SrI LAnkA In SOuth AsIA. (B) ClOsE up OF SrI LAnkA, wIth cOllEctIOn lOcAlItIEs OF Campostichomma And Devendra spEcIEs: whItE squArE, Devendra pardalis (SImOn, 1898), D. pumilus (SImOn, 1898) And D. saama sp. nov.; whItE cIrclE, D seriatus (SImOn, 1898) And Devendra amaiti sp. nov.; blAck squArE, Compostichomma harasbedda sp. nov.; blAck cIrclE, Compostichomma alawala sp. nov.; blAck trIAnglE, Compostichomma mudduk sp. nov. ThE cOllEctIOn dAtA OF Compostichomma manicatum KArsch, 1892 dO nOt prOVIdE A spEcIFIclOcAlIty In SrI LAnkA, thus thE spEcIEs Is nOt shOwn In thIs mAp.