Machimosaurini Jouve et al., 2016

Young, Mark T., Wilberg, Eric W., Johnson, Michela M., Herrera, Yanina, Brandalise, Marco de Andrade, Brignon, Arnaud, Sachs, Sven, Abel, Pascal, Foffa, Davide, Fernández, Marta S., Vignaud, Patrick, Cowgill, Thomas & Brusatte, Stephen L., 2024, The history, systematics, and nomenclature of Thalattosuchia (Archosauria: Crocodylomorpha), Zoological Journal of the Linnean Society 200 (2), pp. 547-617 : 596

Machimosaurini Jouve et al., 2016 (Zoological Code)

Machimosaurini Jouve et al., 2016: 6 (PhyloCode)

RegNum registration number 798.

Type genus: Machimosaurus von Meyer 1837 (emend. von Meyer 1838).

Etymology

‘Pertaining to Machimosaurus ’. The stem Machimosaur - is based on the genus Machimosaurus . - ini is a Latin masculine plural suffix for ‘pertaining to’. Under Article 29.2 of the Zoological Code, the suffix - ini denotes a tribe rank within the family-group.

Geological range

Middle Jurassic (Bathonian) to Early Cretaceous (Hauterivian or possibly late Barremian?) ( Fanti et al. 2016, Jouve et al. 2016, Cortés et al. 2019, Johnson et al. 2020 a, Young and Sachs 2021).

PhyloCode phylogenetic definition

The largest clade within Teleosauroidea containing Machimosaurus hugii ( von Meyer 1837) but not Neosteneosaurus edwardsi ( Eudes-Deslongchamps 1868) . Definition from Johnson et al. (2022b).

Reference phylogeny

Fig. 3 .

Zoological Code diagnosis and PhyloCode diagnostic apomorphies

Teleosauroid crocodylomorphs with the following unique combination of characters (13): expanded network of neurovascular foramina on the premaxillae, maxillae and dentaries (shared with the teleosaurids Aeolodon and Mycterosuchus ; and also seen in extant crocodylians as housing the dome pressure receptors) (86.1); supratemporal fossae are parallelogram in shape (194.5); supratemporal fenestrae at least twice as long as the anterior width (196.1); supratemporal fenestrae at least twice as long as the orbits (shared with Dyrosauridae ) (198.3); deep and pronounced reception pits conspicuous along the anterior and mid-maxilla (596.1); anterior maxillary tooth crowns have blunt and rounded apices (631.1); anterior-middle tooth crowns not recurved (647.0); presence of false ziphodonty across the dentition (652.2); apical tooth crowns have a strongly developed anastomosed enamel ornamentation (shared with the goniopholidids Goniopholis and Anteophthalmosuchus , and the metriorhynchid Torvoneustes ) (663.1); three sacral vertebrae (708.1); sub-rectangular ischial plate (shared with some metriorhynchids) (804.1); tuberosity of the tibia is angled ventrally (824.1); dorsal osteoderm ornamentation composed of pits of highly variable size and length, and on keeled osteoderms the pits can become elongate pits, especially along the lateral margins (840.3).

Composition

Lemmysuchus , Machimosaurus , and Yvridiosuchus .

Figures

Figure 3. The equal weights, no constraints strict consensus topology (C0). Length = 4164, CI = 0.303, RI = 0.831, RC = 0.251, HI = 0.697, number of most parsimonious cladograms hit the hold limit.Phylogeny pruned to highlight Thalattosuchia (see Supporting Information, Appendix S2, Fig. S1 for the entire topology). A, Crocodylomorpha; B, Crocodyliformes; C, Thalattosuchia; D, Neothalattosuchia; E, Teleosauridae; F, Teleosaurinae; G, Aeolodontini; H, Machimosauridae; I, Machimosaurinae; J, Machimosaurini; K, Euthalattosuchia; L, Metriorhynchidae; M, Metriorhynchinae; N, Rhacheosaurini; O, Geosaurinae; P, Geosaurini. Green lines highlight non-neothalattosuchian thalattosuchians, orange lines highlight teleosauroid thalattosuchians, and blue lines highlight metriorhynchoid thalattosuchians.Images from PhyloPics.org, with the teleosauroid by Scott Hartman, and the metriorhynchid by Dmitry Bogdanov (vectorized by T. Michael Keesey), the only change made was to their colouration, and their license is: https://creativecommons.org/licenses/by/3.0/.