Machimosaurinae Jouve et al., 2016

Young, Mark T., Wilberg, Eric W., Johnson, Michela M., Herrera, Yanina, Brandalise, Marco de Andrade, Brignon, Arnaud, Sachs, Sven, Abel, Pascal, Foffa, Davide, Fernández, Marta S., Vignaud, Patrick, Cowgill, Thomas & Brusatte, Stephen L., 2024, The history, systematics, and nomenclature of Thalattosuchia (Archosauria: Crocodylomorpha), Zoological Journal of the Linnean Society 200 (2), pp. 547-617 : 595-596

Machimosaurinae Jouve et al., 2016 (Zoological Code)

Machimosaurinae Johnson et al., 2020a: 118 (PhyloCode)

RegNum registration number 759.

Type genus: Machimosaurus von Meyer 1837 (emend. von Meyer 1838).

Etymology

‘Pertaining to Machimosaurus ’. The stem Machimosaur - is based on the genus Machimosaurus . - inae, is a Latin feminine plural suffix for ‘pertaining to’. Under Article 29.2 of the Zoological Code, the suffix - inae denotes a subfamily rank within the family-group.

Geological range

Middle Jurassic (Bathonian) to Early Cretaceous (Hauterivian or possibly late Barremian?) ( Fanti et al. 2016, Jouve et al. 2016, Cortés et al. 2019, Johnson et al. 2020 a, Young and Sachs 2021).

PhyloCode phylogenetic definition

Thelargestcladewithin Teleosauroidea containing Machimosaurus hugii ( von Meyer 1837) but not Deslongchampsina larteti ( Eudes-Deslongchamps 1866) , Macrospondylus bollensis ( Jäger 1828) , and Charitomenosuchus leedsi ( Andrews 1909) . Definition from Johnson et al. (2022b).

Reference phylogeny

Fig. 3 .

Zoological Code diagnosis and PhyloCode diagnostic apomorphies

Teleosauroid crocodylomorphs with the following unique combination of characters (7): tooth row and occipital condyle aligned in the same plane (shared with the unnamed Chinese teleosaurid and Metriorhynchoidea ) (3.1); ornamentation restricted to the centre of the frontal (shared with the aeolodontin teleosaurids Aeolodon , Bathysuchus and Sericodon ) (22.1); premaxilla–maxilla suture is ‘U’-shaped in dorsal view (shared with the teleosaurid Mystriosaurus ) (132.1); ulna olecranon process greatly expanded (shared with aeolodontin teleosaurids) (768.1); ilium supraacetabular crest shallow and poorly developed (shared with Metriorhynchoidea ) (793.1); ilium postacetabular process expanded into a ‘fan’-shape (795.1); the forelimbs (humerus + ulna) are approximately half the length of the hindlimbs (femur + tibia) at maturity (shared with the hylaeochampsids Pachycheilosuchus and Pietraroiasuchus ) (808.3).

Composition

Andrianavoay , Neosteneosaurus, Proexochokefalos, and Machimosaurini (composed of Lemmysuchus , Machimosaurus , and Yvridiosuchus ).

Comments

Authorship: While the nomen Machimosaurinae was first used by Johnson et al. (2020a: 118), under the Zoological Code Jouve et al. 2016 is the nominal authority. The nominal author of a family-group is the author who first erected a family-group taxon that is valid (in fulfilment of Article 11), and in accordance with the Zoological Code Principle of Coordination applied to family-group names (Article 36.1). However, under the PhyloCode Johnson et al. (2020a) is the nominal authority.

Figures

Figure 3. The equal weights, no constraints strict consensus topology (C0). Length = 4164, CI = 0.303, RI = 0.831, RC = 0.251, HI = 0.697, number of most parsimonious cladograms hit the hold limit.Phylogeny pruned to highlight Thalattosuchia (see Supporting Information, Appendix S2, Fig. S1 for the entire topology). A, Crocodylomorpha; B, Crocodyliformes; C, Thalattosuchia; D, Neothalattosuchia; E, Teleosauridae; F, Teleosaurinae; G, Aeolodontini; H, Machimosauridae; I, Machimosaurinae; J, Machimosaurini; K, Euthalattosuchia; L, Metriorhynchidae; M, Metriorhynchinae; N, Rhacheosaurini; O, Geosaurinae; P, Geosaurini. Green lines highlight non-neothalattosuchian thalattosuchians, orange lines highlight teleosauroid thalattosuchians, and blue lines highlight metriorhynchoid thalattosuchians.Images from PhyloPics.org, with the teleosauroid by Scott Hartman, and the metriorhynchid by Dmitry Bogdanov (vectorized by T. Michael Keesey), the only change made was to their colouration, and their license is: https://creativecommons.org/licenses/by/3.0/.