Pyrgostylus cf. abreui (Peñas & Rolán 2000)

Sacchetti, Claudia, Landau, Bernard & Ávila, Sérgio P., 2023, The Lower Pliocene marine gastropods of Santa Maria Island, Azores: Taxonomy and palaeobiogeographic implications, Zootaxa 5295 (1), pp. 1-150 : 97

Pyrgostylus cf. abreui (Peñas & Rolán 2000)

Plate 6 K 1 -K 2

cf. * Turbonilla abreui Peñas & Rolán 2000: 70 , figs. 40-45.

cf. Turbonilla abreui Peñas & Rolán, 2000 — Peñas et al. 2000: 186, figs. 27A-B.

cf. Pyrgostylus abreui (Peñas & Rolán, 2000) — Landau & Micali 2021: 314, pl. 121, figs. 1-2.

Santa Maria material examined. Maximum height 6.0 mm, width 3.0 mm. DBUA-F 471-2 (3), Ponta do Castelo, Santa Maria Island, Azores, Touril Complex, Lower Pliocene.

Discussion. The specimens at hand are all too incomplete to offer a description and all lack the protoconch, which is of paramount importance in the classification of pyramidellids. Nevertheless, the specimens can be attributed to the genus Pyrgostylus Monterosato, 1884 , based on their very strong axial sculpture with an occasional varix, and the outer lip also thickened by varix. They are most like P. abreui (Peñas & Rolán, 2000) , an extant species endemic to the island of Madeira, in their relatively slender form and very strong axial sculpture. This species was recently also recorded from the Upper Pliocene western Mediterranean of the Estepona Basin, Spain ( Landau & Micali 2021). In view of the incompleteness of the Azorean fossil specimens, we leave them in open nomenclature.

Distribution. Lower Pliocene: Atlantic, Santa Maria Island, Azores (this paper).

Figures

PLATE 1. A. Danilia pterostomus (Bronn, 1861), height 5.0 mm, width 4.0 mm, Forno da Cré (figure taken from Bronn in Hartung, 1861, pl. 19, fig. 2). B1-B2. Emarginula octaviana Coen, 1939, DBUA-F 487-1, height 1.8 mm, diameter 3.0 mm, Ponta do Castelo. C. Bolma cf. rugosa (Linné, 1767), DBUA-F 471-3, diameter 4.1 mm, Ponta do Castelo. D1-D5. Turbo hartungi Bronn 1861, DBUA-F 1295-1, height 9.0 mm, width 8.0 mm, Malbusca (east cave). E1-E3. Nerita cf. funata Dujardin, 1837, DBUA-F 917-1, height 4.0 mm, width 6.0 mm, Ponta do Castelo. F1-F2. Thericium crenulosum (Bronn, 1862), height 10.0 mm, width 6.0 mm, Ponta do Norte lighthouse, LNEG collection (unnumbered). G1-G2. Thericium hartungi (Mayer, 1862), height 29.0 mm, width 13.0 mm, Pinheiros (figure taken from Bronn in Reiss, 1862, pl. 1, fig. 11). H1-H2. Thericium incultum (Mayer, 1862), DBUA-F 1034-2, height 23.0 mm, width 10.0 mm, Macela viewpoint. I1-I2. Thericium incultum (Mayer, 1862), DBUA-F 1027-1, height 29.0 mm, width 11.0 mm, Malbusca II. J1-J2, K1-K2. Bittium miradouroense nov. sp., Holotype DBUAF 1034-5, height 6.8 mm, width 1.9 mm (J); Paratype DBUA-F 1034-6(1), height 5.3 mm, width 1.9 mm (K), Macela viewpoint. L1-L2, M1-M2. Bittium nov. sp., DBUA-F 488-2, height 4.5 mm, width 1.8 mm (L), height 4.2 mm, width 1.7 mm (M), Ponta do Castelo.

PLATE 2. A1-A2. Bittium nanum (Mayer, 1864), DBUA-F 1296-2, height 4.9 mm, width 2.0 mm, Malbusca. B. Tenagodus (Tenagodus) obtusus (Schumacher, 1817), DBUA-F 1283-2, height 12.0 mm, width 11.0 mm, Ponta do Castelo. C. Epitonium sp., DBUA-F 931-1, height 4.0 mm, width 2.8 mm, Pedra-que-pica. D1-D3. Janthina krejcii, SMF10135a, height 25.0 mm, diameter 32.0 mm, Macela viewpoint (pictures from Beu, 2017, fig. 26). E1-E3. Janthina typica (Bronn, 1861), DBUA-F-428, height 19.0 mm, diameter 21.0 mm, Ponta do Castelo, (pictures from Beu, 2017, fig. 25). F.Cheilea equestris (Linnaeus, 1758), DBUA-F 1283-1, diameter 30.0 mm, Ponta do Castelo. G1-G3. Hipponix sulcatus (Borson, 1820), DBUA-F 937-1, height 5.3 mm, diameter 14.0 mm, Ponta do Castelo. H1-H2. Euspira atlantica (Mayer, 1862), height 16.0 mm, diameter 19.0 mm, Ponta do Norte lighthouse, LNEG collection (unnumbered). I. Triphorid s.l., DBUA-F 471-5, height 3.2 mm, width 1.3 mm, Ponta do Castelo. J1-J2. Cerithiopsis cf. tubercularis (Montagu, 1803), DBUA-F 1023-1, height 3.0 mm, width 1.2 mm, Ponta do Castelo. K. Petaloconchus intortus (Lamarck, 1818), DBUA-F 946-2, height 13.0 mm, width 6.0 mm, Ponta dos Cedros. L. Thylacodes arenarius (Linnaeus, 1758), DBUA-F 869-1, height 33.0 mm, width 24.0 mm, Ponta dos Cedros. M1-M3. Alvania cancellata (da Costa, 1778), DBUA-F 1296-3, height 4.0 mm, width 2.2 mm, Malbusca.

PLATE 6. A1-A2. Conus s.l. sp. A, DBUA-F 1292-1, height 19.0 mm, diameter 13.0 mm, Ponta dos Frades. B. Tomellana sp., height 25.0 mm (apical fragment), Ponta do Norte lighthouse, LNEG collection (unnumbered). C, D. Crassopleura maravignae (Bivona, 1838), height 12.0 mm, width 4.5 mm, Pinheiros (figure C taken from Bronn in Reiss, pl. 1, fig. 9 and figure D taken from Mayer, 1864, pl. 6, fig. 50). E1-E2. Psilaxis simplex (Bronn, 1831), DBUA-F 828-3 height 1.5 mm, width 4.5 mm, Ponta dos Frades. F. Mathilda quadricarinata (Brocchi, 1814), DBUA-F 1023-2, height 1.9 mm, width 1.1 mm, Ponta do Castelo. G1-G2. Acteon semistriatus (Férussac, 1822), DBUA-F 191-9, height 7.3 mm width 3.9 mm, Malbusca. H1-H2. Cylichna sp., DBUA-F 1034-1, height 10.0 mm, width 5.0 mm, Macela viewpoint. I1-I3. Roxania utriculus (Brocchi, 1814), DBUA-F 1027-2, height 8.2 mm, width 4.5 mm, Malbusca II. J1-J2. Scaphander lignarius (Linnaeus, 1758), height 27.0 mm, width 15.5 mm, unknown locality, LNEG collection (unnumbered). K1-K2. Pyrgostylus cf. abreui (Peñas & Rolán, 2000), DBUA-F 471-2, height 6.0 mm, width 3.0 mm, Ponta do Castelo. L1-L3. Clathrella clathrata (Philippi, 1844), DBUA-F 471-1, height 6.0 mm, width 7.0 mm, Ponta do Castelo. M, N. Ranella bicoronata Bronn, 1862 species inquirenda, height 19.0 mm, width 15.0 mm, Pinheiros (figure M taken from Bronn in Reiss, 1862, pl. 1, fig. 8 and figure N taken from Mayer, 1864, pl. 7, fig. 53).