Favartia suboblonga ( d’Orbigny, 1852 )

Sacchetti, Claudia, Landau, Bernard & Ávila, Sérgio P., 2023, The Lower Pliocene marine gastropods of Santa Maria Island, Azores: Taxonomy and palaeobiogeographic implications, Zootaxa 5295 (1), pp. 1-150 : 76

Favartia suboblonga ( d’Orbigny, 1852)

Plate 5 G 1 -G 2

Murex oblongus Grateloup 1833 , pl. 100 (non M. oblongus Brocchi, 1814 ).

Murex oblongus Grateloup 1847 , pl. 3, fig. 13 (non Murex oblongus Brocchi, 1814 ).

* Murex suboblongus d’Orbigny 1852: 73 (nom. nov. pro Murex oblongus Grateloup, 1833 , non Brocchi, 1814).

Murex incisus Broderip —Ĥrnes 1856: 223, pl. 23, fig. 7.

Murex incisus Brod. — D’Ancona 1871: 23, pl. 4, fig. 6.

? Murex (Muricidea) kostejanus Boettger 1902: 29 .

Murex (Favartia) incisus Brod. — Sacco 1904: 20, pl. 5, fig. 19-22.

? Muricidea kostejana (Boettger) — Zilch 1934: 252, pl. 15, fig. 81.

Murex (Favartia) excisus var. Peyroti Montanaro 1935: 27, pl. 2. fig. 11.

Aspella (Favartia) incisa excisa Grateloup , sp. 1833—Glibert 1952: 298, pl. 6, fig. 10.

Aspella (Favartia) incisa var. excisa Grateloup, 1833 — Brébion 1964: 377.

Favartia incisa (Broderip, 1832 [sic])— Inzani & Bertarelli 1985 b: 27, pl. 1, fig. 4.

Favartia incisa (Broderip, 1832 [sic])— Andreoli & Marsigli 1992: 7, pl. 2, fig. 4.

Dermomurex scalarioides ( Blainville, 1829) — Cavallo & Repetto 1992: 86, fig. 185.

Favartia cf. incisa (Broderip, 1832 [sic])— Cavallo & Repetto 1992: 90, fig. 194.

Favartia cf. incisa (Broderip, 1832 [sic])—Muñiz-Solís & Guerra-Merchán 1994: 28, figs. 6C-D.

Favartia (Favartia) suboblonga (d’Orbigny) — Vokes 1994: 122, pl. 19, fig. 3.

Aspella (Favartia) incisa (Broderip, 1832 [sic]) — Bałuk 1995: 221, pl. 26, figs. 7-9, pl. 38, fig. 5.

Favartia suboblonga ( D’Orbigny, 1852) — Merle 1999: 300, text-figs. 39, 40, pl. 5, figs. 7-9, pl. 6, figs. 1-4.

Favartia excisa ( Grateloup, 1833) — Chirli 2000: 23, pl. 11, figs. 9-12.

Favartia incisa (Broderip, 1832 [sic])— Chirli 2000: 24, pl. 11, figs. 13-16.

Favartia suboblonga ( d’Orbigny, 1852) — Landau et al. 2007: 46, text-fig. 12, pl. 12, figs. 5-8.

Favartia suboblonga ( d’Orbigny, 1852) — Landau et al. 2013: 161, pl. 24, fig. 5.

Favartia suboblonga ( d’Orbigny, 1852) — Goret et al. 2013: 17, text-fig 13 a, b, pl. 4, fig. 6.

Favartia suboblonga ( d’Orbigny, 1852) — Ceulemans et al. 2016b: 40, pl. 1, figs. 8, 9.

Favartia suboblonga ( d’Orbigny, 1852) —Landau et al. 2019a: 21, pl. 18, figs. 1-3.

non Murex (Favartia) suboblongus D’Orb. —Cossmann & Peyrot (partim) 1924: no. 752, pl. 12, figs. 21-22 [unnamed species fide Merle, 1999].

non Murex (Favartia) suboblongus D’Orb. —Cossmann & Peyrot (partim) 1924: no. 752, pl. 12, fig. 23 [= Pygmaepterys subdecussatus ( D’Orbigny, 1852) ].

non Murex (Favartia) suboblongus var. quadrangulatus Cossmann & Peyrot 1924 : no. 753, pl. 12, figs. 31, 32, 43 [= Favartia excisa ( Grateloup, 1833].

Santa Maria material examined. Maximum height 19.0 mm, width 10.0 mm. DBUA-F 946-3 (1), Ponta dos Cedros, Santa Maria Island, Azores, Touril Complex, Lower Pliocene.

Discussion. Unfortunately, the Azorean specimen is decorticated, giving the surface a somewhat different aspect to that usually attributed to this species. Nevertheless, the size, general shape, penultimate whorl with two strong cords, and the last whorl with strong varices and a long recurved P1 (terminology follows Merle 1999, 2001) at the shoulder are consistent with Favartia suboblonga ( d’Orbigny, 1852) . For detailed description and discussion on the species, see Landau et al. (2009, 2013, 2019).

Distribution. Middle Miocene (Langhian and Serravallian): Atlantic, Loire Basin, France (Glibert 1952), Aquitaine Basin (Cossmann & Peyrot 1924); Paratethys, Austria (Hörnes 1853), Poland ( Bałuk 1995),? Romania ( Boettger 1902; Zilch 1934); Proto-Mediterranean (Serravallian), Karaman Basin ( Landau et al. 2013). Upper Miocene (Tortonian): Atlantic, NW France ( Brébion 1964; Landau et al. 2019a); Proto-Mediterranean, Italy ( Montanaro 1935). Lower Pliocene: Atlantic, NW France ( Brébion 1964; Ceulemans et al. 2016b), Santa Maria Island, Azores (this paper); western Mediterranean, France ( Goret et al. 2013); central Mediterranean, Italy ( Chirli 2000). Upper Pliocene: western Mediterranean, Estepona Basin ( Landau et al. 2007); central Mediterranean, Italy ( Sacco 1904; Cavallo & Repetto 1992; Andreoli & Marsigli 1992). Upper Pliocene-Pleistocene:Atlantic NW France ( Brébion 1964).

Figures

PLATE 1. A. Danilia pterostomus (Bronn, 1861), height 5.0 mm, width 4.0 mm, Forno da Cré (figure taken from Bronn in Hartung, 1861, pl. 19, fig. 2). B1-B2. Emarginula octaviana Coen, 1939, DBUA-F 487-1, height 1.8 mm, diameter 3.0 mm, Ponta do Castelo. C. Bolma cf. rugosa (Linné, 1767), DBUA-F 471-3, diameter 4.1 mm, Ponta do Castelo. D1-D5. Turbo hartungi Bronn 1861, DBUA-F 1295-1, height 9.0 mm, width 8.0 mm, Malbusca (east cave). E1-E3. Nerita cf. funata Dujardin, 1837, DBUA-F 917-1, height 4.0 mm, width 6.0 mm, Ponta do Castelo. F1-F2. Thericium crenulosum (Bronn, 1862), height 10.0 mm, width 6.0 mm, Ponta do Norte lighthouse, LNEG collection (unnumbered). G1-G2. Thericium hartungi (Mayer, 1862), height 29.0 mm, width 13.0 mm, Pinheiros (figure taken from Bronn in Reiss, 1862, pl. 1, fig. 11). H1-H2. Thericium incultum (Mayer, 1862), DBUA-F 1034-2, height 23.0 mm, width 10.0 mm, Macela viewpoint. I1-I2. Thericium incultum (Mayer, 1862), DBUA-F 1027-1, height 29.0 mm, width 11.0 mm, Malbusca II. J1-J2, K1-K2. Bittium miradouroense nov. sp., Holotype DBUAF 1034-5, height 6.8 mm, width 1.9 mm (J); Paratype DBUA-F 1034-6(1), height 5.3 mm, width 1.9 mm (K), Macela viewpoint. L1-L2, M1-M2. Bittium nov. sp., DBUA-F 488-2, height 4.5 mm, width 1.8 mm (L), height 4.2 mm, width 1.7 mm (M), Ponta do Castelo.

PLATE 2. A1-A2. Bittium nanum (Mayer, 1864), DBUA-F 1296-2, height 4.9 mm, width 2.0 mm, Malbusca. B. Tenagodus (Tenagodus) obtusus (Schumacher, 1817), DBUA-F 1283-2, height 12.0 mm, width 11.0 mm, Ponta do Castelo. C. Epitonium sp., DBUA-F 931-1, height 4.0 mm, width 2.8 mm, Pedra-que-pica. D1-D3. Janthina krejcii, SMF10135a, height 25.0 mm, diameter 32.0 mm, Macela viewpoint (pictures from Beu, 2017, fig. 26). E1-E3. Janthina typica (Bronn, 1861), DBUA-F-428, height 19.0 mm, diameter 21.0 mm, Ponta do Castelo, (pictures from Beu, 2017, fig. 25). F.Cheilea equestris (Linnaeus, 1758), DBUA-F 1283-1, diameter 30.0 mm, Ponta do Castelo. G1-G3. Hipponix sulcatus (Borson, 1820), DBUA-F 937-1, height 5.3 mm, diameter 14.0 mm, Ponta do Castelo. H1-H2. Euspira atlantica (Mayer, 1862), height 16.0 mm, diameter 19.0 mm, Ponta do Norte lighthouse, LNEG collection (unnumbered). I. Triphorid s.l., DBUA-F 471-5, height 3.2 mm, width 1.3 mm, Ponta do Castelo. J1-J2. Cerithiopsis cf. tubercularis (Montagu, 1803), DBUA-F 1023-1, height 3.0 mm, width 1.2 mm, Ponta do Castelo. K. Petaloconchus intortus (Lamarck, 1818), DBUA-F 946-2, height 13.0 mm, width 6.0 mm, Ponta dos Cedros. L. Thylacodes arenarius (Linnaeus, 1758), DBUA-F 869-1, height 33.0 mm, width 24.0 mm, Ponta dos Cedros. M1-M3. Alvania cancellata (da Costa, 1778), DBUA-F 1296-3, height 4.0 mm, width 2.2 mm, Malbusca.

PLATE 5. A1-A2. Tritia atlantica (Mayer, 1862), DBUA-F 1295-2, height 13.0 mm, width 6.5 mm, Malbusca (east cave). B1-B2 C1-C2. Tritia serraticosta (Ĥrnes, 1852), DBUA-F 1292-A, height 6.2 mm, width 4.1 mm (B); height 8.1 mm, width 4.2 mm (C), Macela viewpoint. D. Tritia vetulum (Mayer, 1862), height 12.0 mm, width 6.0 mm, Pinheiros (figure taken from Bronn in Reiss, 1862, pl. 1, fig. 5). E1-E2. Aplus sp., height 7.5 mm, width 4.0 mm, Ponta do Castelo, LNEG collection (unnumbered). F1- F2. Orania cf. bellardiana (Foresti, 1884), DBUA-F 828-2, height 17.0 mm, width 12.0 mm, Ponta dos Frades. G1-G2. Favartia suboblonga (d’Orbigny, 1852), DBUA-F 946-3, height 19.0 mm, width 10.0 mm, Ponta dos Cedros. H1-H2. Bellardithala obsoleta (Brocchi, 1814), DBUA-F 542-4, height 5.9 mm, width 2.0 mm, Ponta dos Frades. I1-I2 J. Cancilla volvaria (Bronn, 1862), DBUA-F 937-2, height 10.9 mm, width 4.8 mm (I), Ponta do Castelo; height 15.5 mm, width 5.5 mm (J), Pinheiros (figure taken from Bronn in Reiss, 1862, pl. 1, fig. 4). K. Mitra (s.l.) peregrinula Mayer, 1864 species inquirenda, height 5.5 mm, width 2.5 mm, Feteirinhas (= Ponta das Salinas) (figure taken from Mayer, 1864, pl. 7, fig. 61). L1-L2. Mitra (s.l.) sp., DBUA-F 1295-3, height 12.9 mm, width 7.0 mm, Malbusca (east cave). M. Africonus trochilus (Mayer, 1864), length 16.0 mm, width 9.0 mm (incomplete spire fragment), Pinheiros, (figure taken from Mayer, 1864, pl. 7, fig. 60). N1-N2. Lautoconus cf. anomalomamillus (Sacco, 1893), DBUA-F 924-1, height 22.0 mm, width 11.0 mm, Malbusca. O. Plagioconus sp., height 35.5 mm, width 16.0 mm, unknown locality, LNEG collection (unnumbered). P1-P2 Q. Stephanoconus candidatus (Mayer, 1862), DBUA-F 913, diameter 19.0 mm (apical fragment of juvenile), Ponta dos Frades (P); diameter 40.0 mm, height 22.0 mm (incomplete spire fragment), Pinheiros (figure taken from Bronn in Reiss, pl. 1, fig. 1).