Aplus subpusillus (Hoernes & Auinger, 1890)

Harzhauser, Mathias & Landau, Bernard M., 2024, The Colubrariidae, Eosiphonidae, Melongenidae, Pisaniidae, Prodotiidae and Tudiclidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea, Zootaxa 5427 (1), pp. 1-110 : 34-35

publication ID	https://doi.org/10.11646/zootaxa.5427.1.1
publication LSID	lsid:zoobank.org:pub:923206B0-E8C5-4FD5-B882-55009ABB0282
persistent identifier	https://treatment.plazi.org/id/03CE9F1C-FFBB-0C65-FF65-FE83EF11FAC3
treatment provided by	Plazi (2024-03-19 21:17:48, last updated 2024-03-19 21:37:23)
scientific name	Aplus subpusillus (Hoernes & Auinger, 1890)
status

Treatment

Aplus subpusillus (Hoernes & Auinger, 1890) View in CoL

Figs 15B View FIGURE 15 , 17A–C View FIGURE 17

* Pollia subpusilla nov. form.—Hoernes & Auinger 1890: 242, pl. 28, figs 12–13.

P [ollia] subpusilla — De Gregorio 1885b: 140.

Pollia (Engina) subpusilla Hö. Au. View in CoL — Boettger 1906: 32.

Pollia subpusilla R.H. I Auing. View in CoL — Friedberg 1912: 186, pl. 11, fig. 1.

Cantharus (Pollia) subpusilla Hörnes-Auinger — Csepreghy-Meznerics 1950: 50.

C [antharus]. (P [ollia]). subpusillus (R. Hörn. et Au.) View in CoL — Sieber 1958: 151.

Cantharus (Pollia) lapugyensis (Hoern. & Auing.) — Báldi 1960: 67, pl. 2, fig. 7 [non Aplus lapugyensis (Hoernes et Auinger, 1890) View in CoL ].

Cantharus (Pollia) pusillus subpusillus Hoernes & Auinger, 1891 — Strausz 1966a: 307, fig. 139.

Euthria subpusillus R. Hoernes et Auinger, 1879 — Zelinskaya et al. 1968: 192, pl. 45, figs 22–23.

Cantharus (Pollia) aquitanensis Peyr. — Csepreghy-Meznerics 1969: 85, pl. 3, figs 20–21 [non Aplus aquitanensis ( Peyrot, 1928) View in CoL ].

Cantharus (Pollia) subpusillus Hoernes et Auinger — Csepreghy-Meznerics 1972: 28, pl. 11, fig. 13.

Cantharus (Pollia) aquitanensis Peyr. — Csepreghy-Meznerics 1972: 28, pl. 11, figs 17–18 [non Aplus aquitanensis ( Peyrot, 1928) View in CoL ].

Cantharus subpusillus (Hoernes & Auinger, 1890) — Bałuk 1995: 241, pl. 37, fig. 4.

Aplus subpusillus (Hoernes et Auinger, 1885) View in CoL — Kovács & Vicián 2023: 250, figs 12D–E.

non Cantharus (Pollia) subpusillus R. Hoernes et Auinger, 1890 — Iljina 1993: 96, pl. 13, figs 1–2 [= Aplus sinzovi ( Bidzinashvili, 1975) ].

Type material. Lectotype (designated by De Gregorio 1885b: 140), NHMW 1859 View Materials /0045/0123, SL: 9.3 mm, MD: 5.7 mm, Niederleis ( Austria), illustrated in Hoernes & Auinger (1890: pl. 28, fig. 13), Figs 17A View FIGURE 17 1 –A View FIGURE 1 2 View FIGURE 2 . NHMW 1866 View Materials /0001/1066, SL: 10.8 mm, MD: 6.1 mm, Forchtenau ( Austria), illustrated in Hoernes & Auinger (1890: pl. 28, fig. 12), Figs 17B View FIGURE 17 1 –B View FIGURE 1 2 View FIGURE 2 , 15B View FIGURE 15 . NHMW 1863 View Materials /0015/0465, SL: 11.0 mm, MD: 6.2 mm, Forchtenau ( Austria), Fig. 17C View FIGURE 17 .

Additional paralectotypes. 2 spec., NHMW 1866 View Materials /0001/0849, Niederleis ( Austria) . 35 spec., NHMW 1863 View Materials /0015/0668, Niederleis ( Austria) . 26 spec., NHMW 1869 View Materials /0001/0484, Forchtenau ( Austria) .

Revised description. Small, squat biconic shell of up to five teleoconch whorls; apical angle 55–62°. Protoconch low conical of 2.5 convex whorls, diameter: 700 μm, height: 700 μm. Early teleoconch whorls almost flat-sided, periphery at abapical suture, with narrow, deeply incised suture. Sculpture of three close-set spiral cords strongly swollen almost tubercular over ribs. Single secondary spiral cord intercalated between adapical and mid-whorl primary spirals on third teleoconch whorl. Additional secondary spiral appears on penultimate whorl at abapical suture. Last whorl attaining 65–67% of total height, strongly convex with rounded periphery and conical base, bearing 11 axial ribs and ten elevated, rounded spiral cords; one secondary spiral cord intercalated in each interspace, secondary between two adapical primaries more prominent than others.Aperture moderately wide, ovate. Columella excavated in upper half. Columellar callus forming broad, thickened rim, sharply delimited from base, with about four weak to subobsolete columellar folds, angled at transition to siphonal canal, accentuated by abapical columellar fold. Anal canal broad U-shaped, moderately incised in thickened adapical tip of aperture, accentuated by small parietal denticle and larger anal denticle. Outer lip strongly thickened by terminal varix, with seven elongated denticles almost extending to peristome. Siphonal canal moderately short, wide, slightly deflected to the left, weakly notched.

Discussion. This species is characterized by its small size, squat biconic outline and tubercular sculpture. The superficially similar but larger Aplus moravicus (Hoernes & Auinger, 1890) differs in its convex spire whorls and prominent lirae inside the outer lip. Moreover, it has a longer siphonal canal. This species was confused by Csepreghy-Meznerics (1969, 1972) with Aplus aquitanensis ( Peyrot, 1928) from the Early Miocene of France. Aplus aquitanensis differs from Aplus subpusillus (Hoernes & Auinger, 1890) in its higher last whorl and narrower aperture (see Peyrot 1927: pl. 4, figs 40–41; Lozouet 2021: pl. 40, fig. 9).

Iljina (1993) listed Aplus subpusilla from the Konkian of the Mangyshlak Peninsula ( Kazakhstan). The illustrated Eastern Paratethyan specimens, however, differ considerably from the Central Paratethyan species in their prominent axial ribs, weak spiral sculpture and weaker denticles in the inner lip. This species was described by Bidzinashvili (1975) as Genota sinzovi and therefore, we propose Aplus sinzovi ( Bidzinashvili, 1975) as new combination for it.

Paleoenvironment. Probably inner neritic. At the Niederleis section, sediment and fossils from coastal and lagoonal environments not exceeding 30 m water depth were transported by tempestites into offshore settings ( Mandic et al. 2002).

Distribution in Central Paratethys. Badenian (Middle Miocene): Polish-Ukrainian Fore-Carpathian Basin: Zboriv (= Zborów) ( Ukraine) ( Friedberg 1912); Korytnica Basin: Korytnica ( Poland) ( Bałuk 1995); Vienna Basin: Niederleis ( Austria); Boršov (= Porstendorf), Porzteich at Břeclav ( Czech Republic) (Hoernes & Auinger 1890); Eisenstadt-Sopron Basin: Forchtenau ( Austria) (Hoernes & Auinger 1890); North Alpine-Carpathian Foreland Basin: Lysice ( Czech Republic) (Hoernes & Auinger 1890); Pannonian Basin: Hidas, Letkés, Szokolya ( Hungary) ( Báldi 1960; Strausz 1966a; Kovács & Vicián 2023); Bükk Mountains: Borsodbóta ( Hungary) ( Csepreghy-Meznerics 1972); Făget Basin: CoŞteiu de Sus ( Romania) ( Boettger 1906).

References

Baldi, T. (1960) Tortonische Molluskenfauna von Badener Tegelfazies' aus Szokolya, Nordungarn. Annales historico-naturales Musei nationalis hungarici, pars mineralogica et palaeontologica, 52, 51 - 99. [http: // publication. nhmus. hu / pdf / annHNHM / Annals _ HNHM _ 1960 _ Vol _ 52 _ 51. pdf]

Baluk, W. (1995) Middle Miocene (Badenian) gastropods from Korytnica, Poland; Part II. Acta Geologica Polonica, 45, 153 - 255. [https: // geojournals. pgi. gov. pl / agp / article / view / 13663]

Bidzinashvili, L. M. (1975) NekotoryemaloizuchennyemollyuskisrednegomiotsenaGruzii. Vsesoyuznyinauchno-issledovatel'skii geologorazvedonchnyi neftyanoi institut (VNIGNI), Gruzinskoe otdelenie, Trudy, 188, 99 - 121. [BidZiнasviлi, Л. М. (1975) Нikotoрыi мaлoiZuciнныi мoллюсki срidнigo мiotiнa GрuZii. VсiсoюZный нaucнo-iссЛidovatiЛЬсKiй нiфtЯнoй iнсtitut (VНIGНI), GрuZiнсKoi otdiЛiнii, Tрudы, 188, 99 - 121.]

Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]

Csepreghy-Meznerics, I. (1950) A hidasi (Baranya m.) tortonai fauna. A Magyar Allami Foldtani Intezet Evkonyve, 39, 3 - 106. [https: // epa. oszk. hu / 03200 / 03274 / 00058 / pdf / EPA 03274 _ mafi _ evkonyv _ 1950 _ 02 _ 003 - 103. pdf]

Csepreghy-Meznerics, I. (1969) Nouvelles gasteropodes et lamellibranches pour la faune hongroise des gisements tortoniensinferieurs de la Montagne de Bukk. Annales historico-naturales Musei nationalis hungarici Pars mineralogica et palaeontologica, 61, 63 - 127.

Csepreghy-Meznerics, I. (1972) La faune Tortonienne-inferieure des gisements tufiques de la Montagne de Bukk: Gastropodes I. Az Egri Muzeum Evkonyve (Annales Musei Agriensis), 8, 26 - 46.

De Gregorio, A. (1885 b) Continuazione degli studi su talune conchiglie mediterranee viventi e fossili. Bulletino della Societa Malacologica Italiana, 11, 27 - 203. [http: // biodiversitylibrary. org / page / 39284454]

Friedberg, W. (1912) Mieczaki Miocenskie ziem Polskich. (Mollusca Miocaenica Poloniae). Czesc I. Slimaki i Lodkonogi (Pars I. Gastropoda et Scaphopoda). Muzeum imienia Dzieduszyckich we Lwowie, 2, pp. 113 - 239, pls. 6 - 14. [https: // rcin. org. pl / dlibra / publication / 4795 / edition / 29891 / content]

Hoernes, R. & Auinger, M. (1879 - 1891) Die Gasteropoden der Meeres - Ablagerungen der ersten und zweiten Miocanen Mediterran - Stufe in der Osterreichisch - Ungarischen Monarchie. Abhandlungen der Kaiserlich - Koniglichen Geologischen Reichsanstalt, 12, 1 - 382, 50 pls. [pp. 1 - 52, pls. 1 - 6 (1879), 53 - 112, pls. 7 - 12 (1880), 113 - 152, pls. 13 - 16 (1882), 153 - 192, pls. 17 - 22 (1884), 193 - 232, pls. 23 - 28 (1885), 233 - 282, pls. 29 - 36 (1890), 283 - 330, pls. 37 - 42 (1891), 331 - 382, pls. 43 - 50 (1891), https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405

Iljina, L. B. (1993) Handbook for identification of marine Middle Miocene gastropods of Southwestern Eurasia. Transactions of the Paleontological Instituti, 255, 1 - 151. [Iльiнa, Л. B. (1993) Opрidiлitiль мoрсkik срidнiмiotiнovыk gaсtрopod Юgo-Zapadнoй IvрaZii. Tрudы PaЛioнtoЛogiciсKogo iнсtituta, 255, 1 - 151.]

Kovacs, Z. & Vician, Z. (2023) Buccinoidea (Neogastropoda) assemblage from the Lower Badenian (Middle Miocene) deposits of Letkes (Hungary). Bolletino Malacologico, 59, 222 - 259. https: // doi. org / 10.53559 / BollMalacol. 2023.15

Lozouet, P. (2021) Buccinoidea (Mollusca, Gastropoda, Neogastropoda) de l'Oligocene superieur (Chattien) du bassin de l'Adour (Sud-Ouest de la France). Cossmanniana, 22, 1 - 129.

Mandic, O., Harzhauser, M., Spezzaferri, S. & Zuschin, M. (2002) The paleoenvironment of an early Middle Miocene Paratethys sequence in NE Austria with special emphasis on paleoecology of mollusks and foraminifera. Geobios, Memoire Special, 24, 193 - 206. https: // doi. org / 10.1016 / S 0016 - 6995 (02) 00059 - 1

Peyrot, A. (1928) Conchologie neogenique de l'Aquitaine. Actes de la Societe Linneenne de Bordeaux, 79, 207 - 465, pls. 5 - 14. [https: // www. biodiversitylibrary. org / page / 48879155]

Sieber, R. (1958) Systematische Ubersicht der jungtertiaren Gastropoden des Wiener Beckens. Annalen des Naturhistorischen Museums in Wien, 62, 123 - 192. [https: // opac. geologie. ac. at / ais 312 / dokumente / Annalen _ des _ NHMW _ 1958 _ 062 _ 123 _ 192. pdf]

Strausz, L. (1966 a) Die miozan-mediterranen Gastropoden Ungarns. Akademiai Kiado, Budapest, 693 pp.

Zelinskaya, V. A., Kulichenko, V. G., Makarenko, D. E. & Sorochan, E. A. (1968) Paleontologiceskij spravocnik. Tom 2. Bryukhonogie i lopatonogie mollyuski paleogena i miotsna Ukrainy. Naukova dumka, Kiev, 282 pp. [ZiлiнсkaЯ, V. A., Kuлiciнko, V. G., Мakaрiнko, D. I., & Сoрocaн, I. A. (1968) PaЛioнtoЛogiciсKiй сpрavocнiK. Toм 2. Bрюkoнogii i Лopatoнogii мoЛЛюсKi paЛiogiнa i мiotiнa UKрaiны. Нaukova duмka, Kiiv, 282 pp.]

Figures

FIGURE 1. Measurements for shell morphology; AA: apical angle, SL: shell length, MD: maximum diameter, LWH: last whorl height, AH: aperture height, AL: aperture length, AW: aperture width.

FIGURE 2. Colubraria subobscura (Hoernes & Auinger, 1884). A1–A3. NHMW 1854/0035/0170, paralectotype, Lăpugiu de Sus (Romania). B1–B2. NHMW 1859/0037/0018a, paralectotype, Lăpugiu de Sus (Romania). C1–C3. NHMW 1865/0001/0183, paralectotype, Lăpugiu de Sus (Romania). D1–D2. NHMW 1859/0037/0018b, paralectotype, Lăpugiu de Sus (Romania). E1–E3. NHMW 1874/0025/0021, lectotype, Lăpugiu de Sus (Romania). F1–F2. NHMW 2023/0338/0001, paralectotype, Lăpugiu de Sus (Romania).

FIGURE 15. Protoconchs of Paratethyan Pisaniidae, Prodotiidae and Tudiclidae. A. Aplus moravicus (Hoernes & Auinger, 1890), NHMW 1862/0029/0034, lectotype, Rudice u Blanska (Czech Republic). B. Aplus subpusillus (Hoernes & Auinger, 1890), NHMW 1866/0001/1066, paralectotype, Forchtenau (Austria). C. Aplus transsylvanicus (Hoernes & Auinger, 1890), NHMW 1873/0026/0019b, lectotype. D. Aplus zebus (De Gregorio, 1885a), NHMW 1855/0045/0439, Gainfarn (Austria). E. Aplus anatolicus nov. sp. RGM 784 005, paratype, Seyithasan (Turkey). F. Hilda transsylvanica (Hoernes & Auinger, 1884), NHMW 1876/0011/0023a, lectotype, Lăpugiu de Sus (Romania).G. Janiopsis vindobonensis nov. sp., NHMW 1850/0009/0002, holotype, Baden (Austria). H. Monostiolum hungaricum (Csepreghy-Meznerics, 1954), NHMW 1868/0001/0094, Steinebrunn (Austria). I. Dianthiphos klaudiae (Kovács & Vicián, 2023), NHMW1846/0037/0144a, Steinebrunn (Austria). J. Euthria obelixi nov. sp., NHMW 1865/0015/0060, holotype, Lysice (Czech Republic). K. Tethyspollia mariae (Hoernes & Auinger, 1890), NHMW 1868/0001/0441, paralectotype, Lăpugiu de Sus (Romania).

FIGURE 17. Aplus subpusillus (Hoernes & Auinger, 1890). A1–A2. NHMW 1859/0045/0123, lectotype, Niederleis (Austria). B1–B2. NHMW 1866/0001/1066, paralectotype, Forchtenau (Austria). C. NHMW 1863/0015/0465, paralectotype, Forchtenau (Austria).

Abbreviations

MD	Museum Donaueschingen

Copyright notice

No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.

Taxonomy

Kingdom	Animalia
Phylum	Mollusca
Class	Gastropoda
Order	Neogastropoda
Family	Pisaniidae
Genus	Aplus