Draconarius immensus Xu and Li, 2006

Wang, Xin-Ping & Jäger, Peter, 2010, A review of Coelotinae epigynal teeth morphology, with descriptions of two species from China (Araneae: Amaurobiidae), Journal of Natural History 44 (19 - 20), pp. 1165-1187 : 1182-1184

Draconarius immensus Xu and Li, 2006

( Figures 6–9 )

Material examined

Five males, one female, China: Yunnan, Zhongdian County, 36 km east-southeast Zhongdian , 3500–3550 m, 27°40.9′ N, 100°01.05′ E, overgrown rock hillside with old mixed forest, bamboo, dead wood, vinegar trap, 23–24 August 2003, D.W. Wrase ( SMF) .

Remarks

The male specimens examined are the same as D. immensus but the female differs slightly in the shape and separation of spermathecae. This might be because of the individual variation in vulva structure, which is common in the incertus species group. The female is similar to D. papillatus Xu and Li 2006 , which is also found in Zhongdian County, Yunnan, in addition to its distribution in Sichuan and Tibet (Xu and Li 2006). Draconarius papillatus might be a junior synonym of D. immensus but this needs to be confirmed by collecting and examining male D. papillatus from the holotype locality (Sumdo, 29.1° N, 100.1° E, Xiangcheng County, Sichuan) (Xu and Li 2006). The eyes and body of the specimens examined in this study are also measured for the purpose of future comparison regarding the species status of D. immensus and D. papillatus .

Diagnosis

This species is similar to D. incertus in having a similar epigynum, a bifurcate conductor, and a simple median apophysis, but can be distinguished by the spermathecae extending anteriorly beyond their heads and the spermathecal heads arising medially between spermathecae in the female, and by the short median apophysis, the embolus originating between prolateral and proximal at about the 6 to 7 o’clock position and running more than one-quarter of an oval ( Figures 6 , 7 ).

Description

Female. Large sized Coelotinae , total length 11.50 ( Figure 8 A–C). Dorsal shield of prosoma 5.50 long, 3.16 wide; opisthosoma 6.00 long, 4.20 wide. AME smallest, slightly more than half the size of ALE; ALE largest; posterior eyes subequal, three-quarters the size of ALE (AME 0.13, ALE 0.21, PME 0.15, PLE 0.16); AME separated from each other by their diameter, from ALE by slightly less than AME diameter; PME separated from each other by more than their diameter, widely separated from PLE by slightly more than 1.5 times PME diameter (AME–AME 0.13, AME–ALE 0.10, PME–PME 0.19, PME–PLE 0.25, AME–PME 0.18) ( Figure 8D ). Labium slightly longer than wide (length/width = 1.06) ( Figure 8E ). Promargin of chelicera with three teeth, retromargin with two teeth. Epigynal teeth absent; atrium large, wider than long, anterior atrial margin recurved medially to a distinct septum, lateral atrial margins indistinct, ridges in atrium distinct and extending to anterior half of atrium; copulatory ducts small, originating posteriorly between spermathecae; spermathecae broad, distinctly separated by about half of their width, slightly extending anterior of their heads; spermathecal heads small, arising medially between spermathecae ( Figures 6A,B , 7A,B ).

Male. Large Coelotinae , total length 10.15 ( Figures 8 F–H). Dorsal shield of prosoma 5.15 long, 3.00 wide; opisthosoma 5.00 long, 3.00 wide. AME smallest, ALE largest; posterior eyes subequal in size, slightly smaller than ALE (AME 0.14, ALE 0.18, PME 0.16, PLE 0.17); AME separated from each other by slightly more than half of their diameter, from ALE by one-third of AME diameter; posterior eyes almost equally separated by slightly less than their diameter (AME–AME 0.08, AME–ALE 0.05, PME–PME 0.14, PME–PLE 0.15, AME– PME 0.13) ( Figure 8I ). Labium slightly longer than wide (length/width = 1.16) ( Figure 8J ). Promargin of chelicera with three teeth, retromargin with two teeth. Palpal patellar apophysis short, curved dorsally; RTA more than half of tibial

length, with distinctly protruding distal end; lateral tibial apophysis distinct; cymbial furrow less than half of tibial length; conductor short, with a broad base, a slender, bifurcate apex, a small basal lamella, and a slender dorsal apophysis; median apophysis short, not spoon-shaped; embolus short, filiform, originating between prolateral and proximal at approximately 6 to 7 o’clock position, extending posteriorly to distal part of tibia, anteriorly not coiling beyond distal part of bulb ( Figures 6C,D , 7 C–E).

Relationships

Draconarius immensus could be the sister species to D. incertus .

Distribution

China (Yunnan, Sichuan) ( Figure 9 ) .

Figures

Figure 6. Draconarius immensus Xu and Li 2006: (A,B) female pigynum, ventral and dorsal views; (C,D) male palp, ventral and retrolateral views.

Figure 7. Draconarius immensus Xu and Li 2006: (A,B) female epigynum, ventral and dorsal views; (C–E) male palp, prolateral, ventral and retrolateral views.

Figure 8. Draconarius immensus Xu and Li 2006, female (A–E) and male (F–J): (A–C, F–H) habitus, dorsal, lateral and ventral views; (D,I) eyes, view between front and dorsal; (E,J) labium, ventral view.

Figure 9. Distribution records of Draconarius wrasei sp. nov. (black circle only) and Draconarius immensus Xu and Li 2006 (black square and circle).