Limnogonus (Limnogonus) fossarum fossarum ( Fabricius, 1775 )

Cianferoni, Fabio, Saleem, Rukhshanda, Afsheen, Sumera & Zia, Ahmed, 2021, The water striders of Pakistan (Hemiptera: Heteroptera: Gerridae): new records from Punjab and first checklist for the country, Zootaxa 4996 (3), pp. 469-491 : 473-474

Limnogonus (Limnogonus) fossarum fossarum ( Fabricius, 1775)

Cimex fossarum Fabricius 1775: 727 .

Gerris fossarum: Fabricius 1794: 188 .— Pakistan: Pruthi (1933: 102. Punjab Province, Jhelum District, Chalisa); Khan & Bibi (1999: 102, Punjab Province, Multan District, “aquatic habitats in and around Multan city”).

Hydrometra fossarum: Fabricius 1803: 258 .

Gerris discolor Stål 1859: 265 .

Hydrometra discolor: Mayr 1868: 172 .

Limnogonus fossarum: Stål 1868: 133 .

Limnogonus discolor: Stål 1868: 133 . Syn. Lundblad (1933: 377).

Tenagogonus nymphae Esaki 1925: 58 .

Tenagogonus okinawanus Esaki 1925: 59 . Syn. Andersen (1975: 30).

Limnogonus nymphae: Esaki 1926: 181 . Syn. Esaki (1926: 181).

Limnogonus okinawanus: Lundblad, 1933: 371 .

Limnogonus (Limnogonus) fossarum: Matsuda 1960: 198 .

Material examined. Gujranwala District: Upper Chenab canal, Nandipur, Gujranwala, 32.254099° N 74.269467° E, 30.X.2016, 2 ♀ apt.; idem, 10.VIII.2017, 6 ♂, 7 ♀ macr .; idem, 2 ♂, 2 ♀ macr., det. FC ( CFC) ; idem, 9.X.2017, 3 ♂, 1 ♀ macr .; idem, 29. VI .2018, 3 ♂, 1 ♀ apt.; idem, 14.X.2019, 3 ♂, 2 ♀ macr.; Upper Chenab canal, Master City Housing Scheme, Gujranwala , 32.130067° N 74.214012° E, 30.X.2016, 1 ♂, 3 ♀ apt.; idem, 10.VIII.2017, 2 ♂, 4 ♀ macr.; Upper Chenab canal near Chan Da Qilla , Gujranwala , 32.104081° N 74.190967° E, 30.X.2016, 1 ♂ apt.; idem, 9.X.2017, 1 ♂, 1 ♀ macr .; idem, 10.VIII.2017, 3 ♂, 2 ♀ macr.; Upper Chenab canal, Kamoki-Tatle Aali Road , 31.984152° N 74.170964° E, 30.X.2016, 1 ♂ macr.; idem, 24.XII.2016, 1 ♀ apt .; idem, 9.X.2017, 4 ♂, 2 ♀ macr .; idem, 10.VIII.2017, 3 ♂, 8 ♀ macr.; Nullah Palkhu , Grand Trunk Road Wazirabad , 32.449509° N 74.112278° E, 11. V .2016, 6 ♂, 8 ♀ macr.; idem, 24.IX.2017, 5 ♂, 6 ♀ macr .; idem, 15.X.2017, 2 ♂, 2 ♀ apt., 3 nymphs; Gammon Lake , Wazirabad , 32.465998° N 74.097017° E, 13.XI.2016, 4 ♂, 5 ♀ macr., 1 ♂ apt.; idem, 24.XII.2016, 7 nymphs ; idem, 15.X.2017, 4 ♂, 1 ♀ macr .; idem, 1 ♂, 1 ♀ macr., 1 ♂ apt., det. FC ( CFC); Lower Chenab canal near Fish Farm Chanawan , 32.330808° N 73.888941° E, 24.IX.2017, 1 ♂, 3 ♀ macr.; Nullah near National Highway Kamoki, 31.997910° N 74.217961° E, 30.X.2016, 3 ♂, 5 ♀ macr.; idem, 12.IV.2016, 1 ♂, 1 ♀ macr., 2 ♂, 4 ♀ apt., 2 nymphs ; idem, 1.VIII.2017, 2 ♂ macr., 2 ♂, 2 ♀ apt .; idem, 9.X.2017, 4 ♂, 3 ♀ macr .; idem, 10.VIII.2017, 3 ♂, 8 ♀ macr.; near Noshra Virkan Road , 32.023413° N 74.139566° E, 10.VIII.2017, 4 ♂, 6 ♀ macr. Gujrat District: Upper Jhelum canal near Pathan Colony, Kharian , 32.719544° N 73.795622° E, 17.IX.2017, 2 ♂, 2 ♀ macr.; idem, 22.X.2017, 2 ♂ macr.; Upper Jhelum canal near Simbli Sharif Road Sarai Alamgir , 32.865010° N 73.713452° E, 22.X.2017, 1 ♀ macr.; Halsi Nala, Gujrat, 32.592015° N 74.105765° E, 11. V .2016, 4 ♀ macr. 1 ♂ apt.; idem, 1.I.2017, 4 nymphs ; idem, 17.IX.2017, 10 ♂, 15 ♀ macr .; idem, 22.X.2017, 17 ♂, 13 ♀ macr .; idem, 22.X.2017, 3 ♂, 2 ♀ macr., det. FC ( CFC) ; idem, 30.X.2019, 8 ♂, 6 ♀ macr. Sialkot District: pond near Jaje Village , 32.394221° N 74.628148° E, 29.X.2016, 2 ♂, 3 ♀ macr.; idem, 12.III.2017, 8 nymphs ; idem, 10.VII.2017, 6 ♂, 4 ♀ macr., 7 nymphs ; idem, 2 ♂ apt., det. FC ( CFC); Marala Ravi link canal near Basi wala pul , 32.222449° N 74.590056° E, 12.III.2016, 5 ♂, 3 ♀ macr.; idem, 29.X.2016, 4 ♂, 2 ♀ macr .; idem, 10.VII.2017, 11 ♂, 9 ♀ macr.; Marala Ravi link canal near Kotli Baba Faqir Chand , 32.293986° N 74.526254° E, 12.III.2016, 3 ♂, 2 ♀ macr., 1 nymph; idem, 29.X.2016, 1 ♂, 1 ♀ macr .; idem, 10.VII.2017, 7 ♂, 9 ♀ macr.; pond near Dar Rice Mills , Bhoroki, Pasrur Road , 32.403149° N 74.603264° E, 29.X.2016, 2 ♂, 2 ♀ macr., 1 ♀ apt.; idem, 3.XII.2016, 1 ♀ apt., 2 nymphs ; idem, 16.IX.2017, 4 ♂, 8 ♀ macr .; idem, 7.X.2017, 9 ♂, 10 ♀ macr., 1 ♂, 2 ♀ apt.; Marala Ravi link canal near Canal City Housing Society , 32.475159° N 74.422701° E, 11. V .2016, 6 ♂, 6 ♀ macr., 1 ♂, 1 ♀ apt., 3 nymphs; idem, 28.IX.2017, 1 ♀ macr .; idem, 15.X.2017, 1 ♂, 1 ♀ macr .; idem, 13.X.2019, 4 ♂, 3 ♀ macr.; Upper Chanab canal near Malkhnawala Road , 32.432131° N 74.360571° E, 22.X.2017, 1 ♀ macr.; Nullah near Amrikpura, Sialkot 32.493918° N 74.346205° E, 13.XI.2016, 1 ♂, 6 ♀ macr.; Upper Chanab canal near Zafarwali , 32.478587° N 74.376158° E, 15.X.2017, 2 ♂, 2 ♀ macr.; idem, 13.X.2019, 2 ♂, 5 ♀ macr.; flood drainage canal near Behari Colony , 32.320839° N 74.393796° E, 29.X.2016, 1 ♂, 1 ♀ macr., 1 ♂ apt.; idem, 29.XI.2016, 2 ♂, 2 ♀ macr., 1 ♂, 1 ♀ apt .; idem, 10.VII.2017, 8 ♂, 8 ♀ macr., 2 ♀ apt., 3 nymphs; Marala Ravi link canal near Motra , 32.367782° N 74.419769° E, 13.XI.2016, 9 ♂, 6 ♀ macr.; idem, 24.XII.2016, 9 ♂, 9 ♀ macr .; idem, 10.VIII.2017, 12 ♂, 8 ♀ macr., 3 nymphs; Bambawali Ravi Bedian canal near Circular Road, Daska , 32.328191° N 74.345216° E, 10.VIII.2017, 5 ♂, 2 ♀ macr.

General distribution. The nominate subspecies is known from Pakistan, India, Sri Lanka, Maldives, Myanmar, China (southern and Hainan), Taiwan, Japan (Amami Ôshima and Ryukyu), Thailand, Laos, Cambodia, Vietnam, Malaysia (peninsular and Borneo), Singapore, Indonesia (Sumatra), and Philippines. Other subspecies occur east- ward in the Australasian Region (see Khan & Bibi 1999, Damgaard et al. 2014, Polhemus 2017, Zettel et al. 2017, Kazmi & Ghory 2019).

Remarks. This species was recorded by Pruthi (1933) and Khan & Bibi (1999) from Punjab. The record of Limnogonus sp. by Ahsan & Aziz-Ullah (1974) from Swat (Khyber Pakhtunkhwa Province) probably also refers to this species (see Kazmi & Ghory 2019).