Afroanomalia Varga, 2018

Varga, Oleksandr, 2018, Afroanomalia pseudoclistopyga, a new genus and species of pimpline parasitoid wasp (Hymenoptera: Ichneumonidae: Pimplinae) from the Afrotropical region, Zootaxa 4461 (1), pp. 146-150 : 146-147

Afroanomalia Varga , gen. n.

Type species. Afroanomalia pseudoclistopyga Varga , sp. n.

Composition. The new genus contains only the type species, A. pseudoclistopyga , described below.

Diagnosis. Afroanomalia gen. n. belongs to the tribe Ephialtini among a group of genera comprising Сlistopyga, Sericopimpla , Tromatobia , and Zaglyptus , having a occipital carina distinct and straight dorsally. The newly described genus is apparently closely related to Сlistopyga in having fore wing vein 3 rm absent, hind wing with the distance between distal abscissa of Сu 1 and M longer than vein cu-a, female hypopygium large and projecting, and male malar space emarginated. Afroanomalia gen. n. differs in having the strongly swollen laterally pronotum, wider than mesoscutum in dorsal view, the elongate propodeum, submetapleural carina absent (at least distally), and ovipositor straight, lower valve with strongly backwards projecting apical teeth. The general shape of the apical teeth is similar in Сlistopyga, but no species has these structures as enlarged and projecting.

Description. Head with face flat, about 0.7 × as long as wide; inner margins of eyes weakly emarginate opposite antennal sockets, eyes parallel-sided, not converging ventrally ( Fig. 1b ); clypeus conveX basally, flat apically, about 0.7 × as long as wide, separated from face, apical margin thin and rounded laterally, straight apically; supraclypeal pits deep; malar space about 0.7 × basal width of mandible, emarginated in male ( Fig. 2b ); mandible strongly narrowed apically, twisted, upper tooth longer, than lower tooth; maXillary palps elongate, surpassing ventral part of epicnemial carina; female antenna broken, but has at least 32 flagellomeres (13 flagellomeres on head and 19 on the broken piece glued to a triangle), first flagellomere 1.6 × length of second flagellomere; male antenna with 27 flagellomeres; occipital carina complete, dorsally straight; maXimum diameter of lateral ocellus 0.9 × length of ocellar-ocular distance; temples short and straight, strongly narrowed behind eyes.

Mesosoma. Pronotum strongly swollen dorsally, much wider than mesoscutum in dorsal view ( Fig. 1d ), epomia absent; mesoscutum swollen in lateral view, notauli distinct but weak on basal 0.4; scutellum weakly swollen, with lateral carina present anteriorly; postscutellum trapezoid, mesopleuron with epicnemial carina long, present on lower 0.8 of mesopleuron; mesopleural carina absent ventrally, metapleuron weakly conveX, submetapleural carina absent distally (proXimally not visible in studied specimens, but not forming angle-like projection near middle coXae); pleural carina indistinct; propodeum elongate, without carinae, spiracle round. Legs slender, hind femur 5.2 × as long as wide (6.3 × in male), longer than tibia (not longer than tibia in male); tibial spurs equal length; tarsus long, about 2.0 × longer than hind tibia, third tarsomere about 0.9 × as long as length of tarsomeres 4–5. Female tarsal claws with basal lobes. Fore wing vein 2 rs-m short, about 0.4 × distance between 2 rs-m and 2 m-cu; vein cu-a opposite or basad of Rs&M by about 0.1 of its own length. Hind wing with distance between distal abscissa of Сu 1 and M about 3.8 × longer than vein cu-a. Сu 1 unpigmented (absent in male).

Metasoma generally elongate; first tergite 2.2 × as long as apical width (3.0 × in male), with dorsolateral carina and median longitudinal carina distinct only on anterior 0.2; first sternite long, reaching 0.7 of tergite (0.5 in male); second tergite 1.4 × as long as apical width (2.2 × in male), with dorsolateral grooves, ending close to each other on subapical part of tergite, forming central elliptic flat area; third tergite 1.3 × as long as apical width (1.9 × in male), with the same area, tergites 4–6 with weak dorsolateral swellings; ovipositor straight, length from tip of hypopygium approXimately 2.9 × length of hind tibia, lower valve with swelling on proXimal 0.7, with four apical teeth strongly backwards projecting ( Fig. 1e ).

Etymology. The name consists of two words: «Afro» because the genus is described from the Afrotropical region and «anomalia» because of its unusual morphological characters.

Figures

FIGURE 1. Afroanomalia pseudoclistopyga sp. n., holotype female: a) lateral vieW of habituS; b) frontal vieW of head; c) lateral vieW of head and meSoSoma; d) dorSal vieW of head and meSoSoma (red arroW pointS Strongly SWollen pronotum); e) lateral vieW of the ovipoSitor apex; f) dorSal vieW of metaSomal tergiteS 2–4.

FIGURE 2. Afroanomalia pseudoclistopyga sp. n., paratype male: a) lateral vieW of habituS; b) frontal vieW of head (red arroW pointS malar Space emargination); c) lateral vieW of metaSomal apex and paramereS.