Sinicolinae, Pall-Gergely, 2018

Páll-Gergely, Barna, 2018, Systematic revision of the Plectopylinae (Gastropoda, Pulmonata, Plectopylidae), European Journal of Taxonomy 455, pp. 1-114 : 9

Subfamily Sinicolinae subfam. nov.

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Type genus

Sinicola Gude, 1899 .

Included genera

Gudeodiscus Páll-Gergely, 2013 (with the subgenus Veludiscus Páll-Gergely, 2015), Endothyrella , Halongella Páll-Gergely, 2015 , Sicradiscus Páll-Gergely, 2013 , Sinicola .

The Eocene fossil genus Plectopyloides Yen, 1969 (type species: P. cretaceous Yen, 1969 ; synonyms: Henanspirus Li, 1986 , Menyinspirus Li, 1986 , Yenellus Li, 1986 ) is also classified in this subfamily, because it is most similar to Gudeodiscus (see Páll-Gergely & Asami 2014).

Diagnosis

Shell usually dextral (although most species of Endothyrella are sinistral), flat to conical-globular; umbilicus wide to narrow; protoconch regularly ribbed without any additional sculptural elements (very rarely smooth, see Páll-Gergely et al. 2015 b, 2015 c); apertural fold present or absent; main plica usually absent (present only in Gudeodiscus longiplica Páll-Gergely & Asami, 2016 ). Penis well-developed; penial caecum well-developed to absent; epiphallus present; diverticulum as long as bursa copulatrix (absent in some species, although possibly overlooked during dissection).

Distribution

Same as in the family Plectopylidae , except Thailand and the Malay Peninsula ( Fig. 3 ).

Remarks

The subdivision of the family into two reciprocally monophyletic groups is supported by molecular phylogenetic analysis, which included nearly all genera. Discussion of the phylogenetic results is beyond the scope of the present revision, and will be discussed in a separate paper.

Figures

Fig. 3. Distribution of the extant Sinicolinae subfam. nov. (ruled area) and Plectopylinae Möllendorff, 1898 (grey shaded area). Mainly after Páll-Gergely & Hunyadi (2013) and Páll-Gergely et al. (2015b, 2015c, 2016).