Hoplopholcus bursa, Huber, 2020

Huber, Bernhard A., 2020, Revision of the spider genus Hoplopholcus Kulczyński (Araneae, Pholcidae), Zootaxa 4726 (1), pp. 1-94 : 75-77

Hoplopholcus bursa sp. n.

Figs 372–387

Diagnosis. Easily distinguished from known congeners by large dorsal process of procursus (arrow in Fig. 379 ); also by ventral ‘knee’ of procursus slightly shifted toward retrolaterally, procursus tip with almost straight ventral spine, pointed prolateral process, and additional rounded process prolateral of transparent process; genital bulb with small prolateral process between ventral sclerite and embolar sclerite (arrow in Fig. 384 ). Females are difficult to distinguish externally from congeners; internal median pouch of uterus externus, sclerotized arc, and pair of lateral pouches usually visible in uncleared specimens ( Fig. 375 ); sclerotized ventral arc with strong median sclerite (arrow in Fig. 386 ) (similar to H. trakyaensis ); lateral pouches strongly developed, connected anteriorly to ventral arc, with shallow pockets.

Type material. TURKEY, Bursa: ♂ holotype, ZFMK (Ar 20967), Nilüfer District , [near Ayvaköy] (40.1244°N, 28.7018°E), 570 m a.s.l., 2.vi.2009 (Y.M. Marusik) .

Other material examined. TURKEY, Bursa: 3♂ 1♀ 5 juvs, ZFMK (Ar 20968), same data as holotype . 1♂ 3 juvs, MHNG, “Bursa, 400, s. pierres, 61”, no further data . 1♀ 1 juv., ZFMK (Ar 20969), Uludağ Millî Parkı (Uludağ

National Park) (40.116°N, 29.055°E), 650 m a.s.l., 1 & 4.vi.2009 (Y.M. Marusik). 4♂ 8♀ 8 juvs (2 vials), ZFMK ( Ar 20970–71), Inegöl District, Great Oylat Cave (39.943°N, 29.592°E), in cave, 520 m a.s.l., 3.vi.2009 (Y.M. Marusik) ; 1♀ 1 juv., ZFMK (Ar 21340), same data but outside cave .

Etymology. The species name is derived from the type locality; noun in apposition.

Description. Male (holotype). MEASUREMENTS. Total length 5.6, carapace width 2.1. Distance PME-PME 150 µm; diameter PME 140 µm; distance PME-ALE 50 µm; diameter AME 80 µm; distance AME-AME 50 µm. Leg 1: 47.8 (12.8 + 0.9 + 13.1 + 17.9 + 3.1), tibia 2: 9.2, tibia 3: 7.1, tibia 4: 8.1; tibia 1 L/d: 62.

COLOR (in ethanol). Carapace ochre-yellow, ocular area and clypeus light brown; sternum brown, medially with dark brown radial marks; legs ochre-yellow to light brown, with indistinct darker rings subdistally on femora and tibiae; abdomen gray, with pairs of black marks dorsally in posterior third; ventrally with large light brown mark anteriorly, black mark at spinnerets, and smaller diffuse brown mark in-between.

BODY. Habitus as in H. forskali (cf. Figs 194–195 ). Ocular area slightly elevated. Deep thoracic pit and pair of furrows diverging from pit toward posterior margin. Clypeus unmodified. Sternum wider than long (1.5/0.9), unmodified. Abdomen oval, dorso-posteriorly rounded.

CHELICERAE. As in Figs 380–381 , with pair of latero-distal apophyses provided with two and three modified cone-shaped hairs, respectively; stridulatory ridges fine, barely visible in dissecting microscope.

PALPS. As in Figs 372–374 ; coxa with retrolateral bulge, trochanter barely protruding ventrally, femur with distinct dark line retrolaterally and prolateral stridulatory pick; procursus ( Figs 378–379 ) with small but distinct ventral ‘knee’ slightly shifted toward retrolaterally, with distinctive dorsal process and almost straight ventral spine, with pointed prolateral process and additional rounded process prolateral of transparent process; genital bulb ( Figs 382–385 ) with small prolateral process between ventral sclerite and embolar sclerite (arrow in Fig. 384 ), with distally slightly widening ventral sclerite, with low dorsal membranous process ( Fig. 383 ).

LEGS. Femur 1 with single row of ~30 ventral spines, femur 2 only distally with a few slightly stronger hairs; without curved hairs; few vertical hairs; retrolateral trichobothrium of tibia 1 at 4%; prolateral trichobothrium present on all leg tibiae; tarsi without distinct pseudosegments but with many small platelets.

Male (variation). Tibia 1 in seven other males: 10.8–13.3 (mean 12.0). Modified hairs on cheliceral apophyses slightly variable: of nine males, six were symmetric with two hairs on each side (2+2), three asymmetric 2+3. Number of spines on femur 1 slightly variable. Curved hairs not seen in males but most males with many hairs lost.

Female. In general similar to male but without spines on legs; one female with distinct rows of curved hairs ventrally on tibia 1 and metatarsus 1 (most hairs lost on legs of other females). Tibia 1 in nine females: 7.1–12.1 (mean 10.8).

FEMALE GENITALIA. Epigynum as in Fig. 375 , posterior margin of epigynal plate medially whitish; internal sclerotized arcs, median pouch of uterus externus, and lateral pouches visible in uncleared specimens; with pair of light brown weakly bulging areas in front of epigynum; posterior plate simple, short but wide. Internal genitalia with narrow and rounded median pouch of uterus externus, ventral part of sclerotized arc with strong median sclerite (arrow in Fig. 386 ), lateral pouches strongly developed, connected anteriorly to ventral arc, with shallow pockets; pore plates large, roundish ( Fig. 387 ).

Distribution. Known from several localities in Bursa Province, Turkey ( Fig. 448 ).

Figures

FIGURE 448. Known distributions of Hoplopholcus trakyaensis and H. bursa. TL = type localities.

FIGURES 372–377. Hoplopholcus bursa sp. n. from Turkey, Great Oylat Cave (ZFMK Ar 20970–71). 372–374. Left male palp in prolateral, dorsal, and retrolateral views. 375–377. Epigynum, ventral view, and cleared female genitalia, ventral and dorsal views.

FIGURES 378–387. Hoplopholcus bursa sp. n. from Turkey, Great Oylat Cave (ZFMK Ar 20970–71). 378–379. Left procursus, prolateral and retrolateral views (arrow points at distinctive dorsal process). 380–381. Male chelicerae, frontal and lateral views. 382–385. Left genital bulb, dorsal, prolateral, ventral, and retrolateral views (arrow points at distinctive prolateral process). 386–387. Cleared female genitalia, ventral and dorsal views (arrow points at distinctive median sclerite of ventral arc). mp, median pouch of uterus externus; lp, lateral pouch of uterus externus. Scale lines: 0.5 mm.

FIGURES 194–199. Hoplopholcus spp., live specimens. 194–197. H. forskali (Thorell, 1871), males from Taşkent, females from Zindan Mağarası (before and after egg-laying). 198. H. konya sp. n., female from Körükini Mağarası. 199. H. longipes (Spassky, 1934), male from Kintrishi Protected Area.