Purusha paradoxa ( Gerstaecker, 1895 )

Constant, Jérôme, 2020, Revision of the Eurybrachidae (XV). The Oriental genus Purusha Distant, 1906 with two new species and a key to the genera of Eurybrachini (Hemiptera: Fulgoromorpha: Eurybrachidae), European Journal of Taxonomy 602, pp. 1-40 : 13-18

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https://doi.org/ 10.5852/ejt.2020.602

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scientific name

Purusha paradoxa ( Gerstaecker, 1895 )


Purusha paradoxa ( Gerstaecker, 1895) View in CoL

Figs 2 View Fig , 4–8 View Fig View Fig View Fig View Fig View Fig

Messena View in CoL (?) paradoxa Gerstaecker, 1895: 33 View in CoL [described; allied to Purusha reversa View in CoL ].

Purusha paradoxa – Distant 1906b: 204 View in CoL [transferred to Purusha View in CoL ]. — Metcalf 1956: 8 [catalogued].



Easily recognized by the following combination of characters: (1) tegmina with numerous small, round, black spots on membrane, more or less arranged in rows parallel to apical margin ( Fig. 6A View Fig ); (2) posterior wings brown with numerous small, black-brown spots on apical half ( Fig. 6 View Fig A–B); (3) ventral margin of gonostyli with internobasal process projecting posteriorly ( Fig. 8B View Fig ); (4) anterolateral spine of phallus much smaller than posterolateral one ( Fig. 8F View Fig ); (5) head, pro- and mesonotum brown, concolorous ( Fig. 6C View Fig ).


Immediately recognized by the combination of the following characters: (1) tegmina with a conspicuous white waxy spot at half length, not touching the costal margin, and numerous minute, round, black spots on the membrane, arranged in 2–3 rows parallel to apical margin ( Figs 5A View Fig , 7A View Fig ); (2) posterior wings with numerous brown spots, sometimes merging together, more or less arranged in rows parallel to apical margin ( Figs 5A View Fig , 7A View Fig ); (3) head, pro- and mesonotum brown, concolorous ( Figs 5C View Fig , 7C View Fig ).

Differential diagnosis


Easily separated from males of P. pulverosa and P. vietnamica sp. nov. (and probably P. bellissima sp. nov.) by the combination of characters (1) and (2) of diagnosis. From P. reversa , it is better separated based on characters (3)–(4) of diagnosis: (3) ventral margin of gonostyli without internobasal process projecting posteriorly in P. reversa ( Fig. 17B View Fig ); (4) anterolateral spine of phallus of equal size in P. reversa ( Fig. 17G View Fig ).


The specific name is derived from the Latin adfjective ‘ paradoxa ’, meaning ‘paradoxical’.

Material examined

Holotype ( Fig. 5 View Fig )

INDONESIA • ♀; Java; “ Paradoxa ”, “Zool. Mus. Greifswald, II 27390”, “ Typus Messena paradoxa ”; ZIMG. In the original description, Gerstaecker (1895) stated that the specimen was collected by Hans Fruhstorfer (1866–1922) in the mountains of West Java.

Additional material

INDONESIA – Java • 1 ♀; “Tjibarangbang” [ Cibaregbeg ]; [6°50′ S, 106°39′ E]; 1939; Mrs M.E. Walsh leg.; ZML GoogleMaps 1 ♀; Sukabumi ; [6°55′ S, 106°56′ E]; 1893; H. Fruhstorfer leg.; MMBC GoogleMaps 1 ♀; Sukabumi ; [6°55′ S, 106°56′ E]; alt. 2000 ft; 8 Jan. 1940; J.M.A. van Groenendael leg.; RMNH GoogleMaps 1 ♀; Sukabumi , “Djampang Tengah” [Jampangtengah]; [7°03′ S, 106°48′ E]; alt. 18–2200 ft; 8 May 1939; J.M.A. van Groenendael leg.; RMNH GoogleMaps 1 ♀; “Soekaboemi” [ Sukabumi ]; [6°55′ S, 106°56′ E]; Ouwens leg.; ZMPA GoogleMaps 1 ♀; Staudinger leg.; NHRS . – Sumatra • 1 ♂, 1 ♀; Muller leg.; RMNH 1 ♂; “Sum” [Sumatra], “Serapai Kor.” [ Korintji ]; [1°41′ S, 101°15′ E]; 1915; Edw. Jacobson leg.; RMNH GoogleMaps 1 ♀; Sumatra , Excell. v. Studt G. leg.; MFNB .

Measurements and ratios

♂: LT (n = 2): 23.8 mm (23.5–24.2); LTg/BTg = 2.3; BV/LV = 3.8; BF/LF = 1.6; LP+LM/BT = 0.68. ♀: LT (n = 5): 28.7 mm (27.3–31.4); LTg/BTg = 1.9–2.1; BV/LV = 4.3; BF/LF = 1.5; LP+LM/BT = 0.73.

Supplementary description

Male genitalia ( Fig. 8 View Fig )

Pygofer with lateral portion of posterior margin strongly projecting in a laminate process directed posterodorsally in lateral view and slightly laterally in ventral view; process narrowly rounded apically; ventral half of posterior margin excavate in lateral view; ventral portion of posterior margin bisinuate in ventral view; dorsal portion of pygofer strongly narrowing with posterior margin strongly excavate; anterior margin sinuate in lateral view ( Fig. 8 View Fig A–B). Anal tube large, broadly obovate, dorsoventrally flattened; apical margin slightly emarginate in dorsal view; lateral margin bisinuate, very broadly rounded on distal half; anal column at basal ¼ ( Fig. 8A, D View Fig ). Gonostyli subrectangular in lateral view, with short apicoventral process rounded apically; dorsal margin with laminate process projecting medially and armed with one tooth at medioanterior angle, two teeth on posterior margin and one strong hook laterally; ventral margin deeply emarginate on distal ⅔, with mediobasal process projecting posteriorly, leaving large central opening in ventral view ( Fig. 8 View Fig A–C). Phallobase robust, about as long as broad in dorsal view, with basolateral process directed posterolaterally and elongate, apical, spinose process ventrally on each side directed posterolaterally; slightly mediodorsally to spinose process, slightly shorter, apically blunt process directed posteriorly ( Fig. 8 View Fig E–F). Phallus with very complicated set of intricate processes: basal strong spine directed laterally; trispinose lateral process with basidorsal spine elongate and curved mediodorsally, posteroventral spine very elongate, sword shaped and curved dorsally and slightly anteriorly on distal portion, and apicodorsal spine incurving, sinuate, projecting mediodorsally and slightly anteriorly on distal portion; mediodorsally, pair of laterally compressed processes higher than long in lateral view and with lateral ridged process ( Fig. 8 View Fig E–F).


Indonesia in the islands of Java and Sumatra ( Fig. 4 View Fig ).


Unknown. The species has not been collected or documented in any way since 1940.


Zoologisches Institut und Museum Greifswald


Moravske Muzeum [Moravian Museum]


National Museum of Natural History, Naturalis


Swedish Museum of Natural History, Entomology Collections


Museo Friulano di Storia Naturale














Purusha paradoxa ( Gerstaecker, 1895 )

Constant, Jérôme 2020

Purusha paradoxa – Distant 1906b: 204

Metcalf Z. P. 1956: 8
Distant W. L. 1906: 204


Gerstaecker C. E. A. 1895: 33
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