Glyphobythus anatolicus Hlaváč & Brachat, 2015

Hlaváč, Peter & Brachat, Volker, 2015, New species of the genus Glyphobythus Raffray, 1904 (Coleoptera: Staphylinidae: Pselaphinae) from Turkey, Zootaxa 3904 (4), pp. 596-599 : 597

Glyphobythus anatolicus Hlaváč & Brachat sp. nov.

( Figs. 1–5 )

Etymology: Named after Anatolia, a historic name for Turkey, the country where the new species has been discovered. Material studied: HOLOTYPE, 1 male: N 41o13' 20 E 31o53' 6, TR Zonguldak 3.5.2014, Devrek., 590 m, Brachat & Meybohm (NMPC). PARATYPES, 32 males, 11 females: same data as holotype (OUMNH, PHC, VBC).

Description. Male: Body ( Fig. 1 ) glossy, dark reddish-brown, legs, antennae and maxillary palpi yellowish-brown, length 1.10–1.15 mm, maximum width of elytra about 0.55 mm. Head and pronotum with sparse, long and uneven pubescence, pubescence on elytra and abdominal tergites even and denser. Head slightly wider than long in both sexes, triangular, vertex about twice as wide as frons measured across supra-antennal prominent tubercles, rostrum with a large deep sulcus reaching vertexal foveae which are located in line of anterior margin of eyes, distance between vertexal foveae slightly wider than distance from fovea to lateral margin of head, vertex with median keel reaching posterior margin of vertexal foveae. Maxillary palpi long, slightly shorter than antennae, palpomere II long, pipe-shaped, palpomere III short, both granulate on dorsal surface, terminal palpomere subtriangular, with well-defined basal setose fovea, 0.20–0.21 mm long. Antennae short, scape very long, 3.75 times as long as wide, slightly expanded from base to apex, three times as long as pedicel, antennomere III elongate, slightly pedunculate, 1.4 times as long as IV, antennomeres IV–VIII subequal in size, spherical, IX and X transverse, IX twice as wide as long, X 2.4 times as wide as long, terminal antennomere 1.8 as long as wide, pointed, relative length of antennomeres: 15.0 / 5.0 / 3.5 / 2.5 / 3.0 / 2.0 / 2.0 / 2.0 / 2.0 / 2.5 / 14.0. Pronotum slightly wider than long and about 1.2 times as long as head, widest in anterior third, lateral foveae well-defined, connected by shallow, well-defined antebasal sulcus, median antebasal fovea absent. Elytra about as long as wide, about 1.8 times as long as pronotum, with two large deep basal setose foveae and well defined entire sutural stria on each elytron. Visible part of abdomen slightly narrower and much shorter than elytra, length of visible abdominal tergites 0.3–0.4 of elytral length, first visible tergite (IV) slightly longer than second (V). Aedeagus ( Figs. 3, 4 ) elongate, 0.45 mm long, median lobe almost symmetrical, its apex bifurcate, parameres curved outwards at apices, with two preapical setae, internal sac bearing four accessory sclerites.

Sexual dimorphism: Females ( Fig. 2 ) have narrower head and much smaller eyes, there is no difference in size of both sexes.

Biology. Females of Glyphobythus have been exclusively found in caves while males can also be found outside, living in leaf litter or under deeply embedded stones ( Jeannel 1950). All specimens of G. anatolicus were collected in a small area at the base of a rocky vertical limestone face ( Fig. 5 ) by sifting mainly Fagus leaf litter.

Differential diagnosis. The new species from Turkey is assigned to the genus Glyphobythus basing on the presence of a well-defined dorsal basal fovea on the palpomere IV in males. The new species differs from all other known Glyphobythus in having this fovea located close to the basal margin of palpomere, far from the apex, in unmodified fore tibiae in both sexes, and in a different shape of the aedeagus.

Distribution. Known only from the type locality, the province Zonguldak of Turkey.

Figures

FIGURE 5. Macrohabitat of the type locality of Glyphobythus anatolicus sp. nov.

FIGURES 1 – 4. 1, Glyphobythus anatolicus sp. nov., male holotype, dorsal habitus (scale = 0.5 mm); 2, female, paratype, dorsal habitus (scale = 0.5 mm); 3, aedeagus, dorsal view (scale = 0.2 mm); 4, aedeagus, lateral view (scale = 0.2 mm).