Tominemoura, Sivec & Stark, 2009

Sivec, Ignac & Stark, Bill P., 2009, Tominemoura, A New Stonefly Genus From Sabah, East Malaysia (Plecoptera: Nemouridae), Illiesia 5 (20), pp. 199-203 : 199-201

publication ID

https://doi.org/ 10.5281/zenodo.4753151

DOI

https://doi.org/10.5281/zenodo.4755691

persistent identifier

https://treatment.plazi.org/id/03CB8799-FFDF-FFB2-FED3-DC056326F8AC

treatment provided by

Felipe

scientific name

Tominemoura
status

gen. nov.

Tominemoura View in CoL , gen. nov.

Type species. Tominemoura trilari View in CoL , sp. nov., by monotypy.

Adult habitus. General dorsal color brown. Head almost entirely dark brown, pronotum brown but with pale areas along lateral and posterior margins ( Fig. 1 View Figs ). Legs banded; hind legs with dark coxa and trochanter; femora with narrow basal and wider median and apical brown bands separated by a wide sub-basal and narrow subapical yellow band. Wing membrane pale brown, veins dark brown.

Gills. Four slender cervical gills present ( Fig. 6 View Figs ); outer gills 3-branched, inner gills 2-branched; all gill rami subequal in length and with common origin on respective inner or outer gill trunk.

Male. Most of dorsal epiproct sclerite membranous but covered with rows of tiny scale-like, triangular spines ( Figs. 2-3 View Figs ). Ventral epiproct sclerite large basally, curved upward as a pair of narrow straps dividing dorsal sclerite and extending around epiproct apex ( Figs. 2, 3, 5 View Figs ); center of ventral sclerite membranous and armed with a triangular patch of stout, backward directed spines visible in lateral aspect ( Fig. 5 View Figs ). Paraprocts with three lobes; outer lobe dorsad to mesal lobe in caudoventral aspect ( Fig. 4 View Figs ), hooked and armed with an apical patch of spines (Fig. 7). Mesal lobe broad at base and strongly narrowed to tips ( Figs. 4 View Figs , 7); inner lobes narrow, weakly sclerotized, much shorter than outer and middle lobes, and without spines. Hypoproct strongly tapered to a broadly triangular process ( Fig. 4 View Figs ) which terminates in a slender upturned filament-like structure not shown in figures; vesicle very long and with parallel margins beyond base ( Fig. 4 View Figs ). Cerci a single weakly sclerotized and somewhat barrel-shaped segment ( Figs. 2-4 View Figs ).

Female. Known from two pharate larvae. Sternum 8 bears a deep mesal notch and sternum 7 has a mesal plate covering at least the base of sternum 8 ( Fig. 15 View Fig ).

Larva. General color pale brown with irregular brown occipital maculations ( Fig. 8 View Figs ); mesonotum and metanotum with pale mesobasal spots. Marginal pronotal bristles long around anterolateral angles, short laterally but with a few longer ones at posterior angles ( Fig. 9 View Figs ); bristles becoming obsolete near median suture along both margins. Mouthparts of typical nemourid type but laciniae terminating in a vertical truncate blade like structure armed along ventroapical margin with several rounded teeth, subequal in size ( Figs. 13-14 View Figs ). Apical maxillary palpal segment bearing a small nipple. Gills as described above ( Fig. 6 View Figs ).

Diagnosis. Adults and larvae are distinguished from other Nemouridae by their unique gill structure which is most similar to Nanonemoura among known genera ( Baumann & Fiala 2001). However, in Tominemoura the respective inner and outer gill rami arise from a common trunk rather than from an elongate major ramus. The gill structure and the 3- lobed paraprocts of Tominemoura suggest it should be placed in subfamily Amphinemurinae (Baumann 1975) where its nearest relative may be Amphinemura .

Distribution. Known from Mt. Kinabalu at sites above 3000 meters.

Etymology. The prefix “ Tomi ” honors our colleague Dr. Tomi Trilar, from the Slovenian Museum of Natural History who provided the first specimens of this interesting stonefly.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Plecoptera

Family

Nemouridae

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