Chaetocnema (Udorpes) modesta Gressitt & Kimoto, 1963

Ruan, Yongying, Yang, Xingke, Konstantinov, Alexander S., Prathapan, Kaniyarikkal D. & Zhang, Mengna, 2019, Revision of the Oriental Chaetocnema species (Coleoptera, Chrysomelidae, Galerucinae, Alticini), Zootaxa 4699 (1), pp. 1-206 : 173-175

72. Chaetocnema (Udorpes) modesta Gressitt & Kimoto, 1963

( Fig. 100 )

Chaetocnema modesta Gressitt & Kimoto, 1963: 780 . TL: China, Hubei (Hupeh). TD: CASC

Distribution: China (Hubei, Sichuan), Vietnam, India.

Host plants: Unknown.

Description: Body length: 1.90–2.10 mm. Body width: 1.15–1.20 mm. Ratio of length of antenna to length of body: 0.70–0.75 mm. Ratio of elytron length (along suture) to width (maximum): 2.60–2.65. Ratio of pronotum width (at base) to length: 1.40–1.45. Ratio of length of elytron to length of pronotum (along middle): 2.65–2.70. Ratio of width of elytra at base (in middle of humeral calli) to width of pronotum at base: 1.15–1.25.

Dorsum metallic blue to coppery. Antennomeres 1–3 completely yellow brown, 5–11 brown to deep brown, apical ones darkened. Tarsi and tibiae yellow brown. Pro- and mesofemur yellow brown. Metafemur dark brown with coppery luster.

Head hypognathous. Frontal ridge narrow and flat. Frons covered with closely placed punctures. Ratio of width of frontal ridge to width of antennal socket: 0.85–1.00. Frontolateral sulcus absent. Head entirely covered with extremely close punctures. Orbit and frons closely covered with white setae. Vertex flat, situated on same level as orbit.

Base of pronotum without short longitudinal impressions. Pronotal base evenly convex. Deep row of large punctures at base of pronotum absent. Area adjacent to mid-basal margin of pronotum covered with punctures. Anterolateral prothoracic callosity weak, truncate and oblique. Lateral margins of pronotum slightly convex, with maximum width situated at apical third. Posterolateral prothoracic callosity poorly developed. Diameter of pronotal punctures as large as distance between them.

Elytra with convex lateral sides. Humeral callus well developed. Scutellar row of elytral punctures geminate and confused. Second to eleventh rows of elytral punctures regular.

First protarsomere slightly larger in male. Excavation on metatibia shallow. Large lateral denticle on metatibia obtuse. Metatibial serration proximal to large lateral denticle absent.

In ventral view, aedeagus parallel-sided, apex narrowing abruptly. Ventral longitudinal groove of aedeagus well developed. Ventral longitudinal ridge in middle of aedeagus absent. Apical denticle of aedeagus in ventral view well-differentiated, long, apically truncate. Minute, transverse wrinkles present throughout ventral side of aedeagus. Aedeagus in lateral view curved at base and middle, straight at apex.

Spermathecal pump much shorter than receptacle, with apex flattened. Vaginal palpus slightly narrowing from base till middle, rest of it approximately parallel-sided. Posterior sclerotization longer than wide. Posterior sclerotization wider than anterior.

Types: Holotype (female, CASC): 1) Suisapa, 1000 M., Lichuan Distr. , W. Hupeh, China, 31.VIII.1948, 2) Gressi, Djou Collrs., 3 ) Holotype Female, Chaetocnema (Chaetocnema) modesta Gressitt & Kimoto [Red Label], 4) California Academy of Sciences , Type No. 13258, 5) 22 Konstantinov, 6) 013 .

Paratypes. 1♂ ♀ ( BMNH):1) Suisapa , 1000 M., Lichuan Distr., W. Hupeh, China, VII–23–48, 2) J. L. Gressi Collector, 3 ) Paratype, Chaetocnema (Chaetocnema) modesta Gressitt & Kimoto. 1♂, KUZC: 1) W. Hupeh, China, Lichuan , Hsiaobo, VIII–8–1948. 2) Gressitt & Djou Collrg; 3 ) Paratype, Chaetocnema modesta n Gressitt & Kimoto. 2 CASC: 1) Suisapa, Lichuan Dist. , W. Rupah (1000 m.), August 26, 1943, leg. Gressi and Djou. 1 CASC: 1) Suisapa, 1000 M., Lichuan Distr., W. Hupeh, China, VII–23–48, 2) J. L. Gressi Collector, 3 ) Paratype, Chaetocnema (Chaetocnema) modesta Gressi & Kimoto. 1♂, CASC: 1) Szechuan , China, NE. of Motauchi, Wanhsien. IX.26.48, 4200–4800 ft., 2) Gressi & Djou Collrs., 3 ) Paratype, Chaetocnema (Chaetocnema) modesta Gressitt & Kimoto.

Material: 1♂ ( IZCAS), China, E-bian, Sichuan ; 1♂ 1♀ ( IZCAS), China, E-mei Mountain , Sichuan ; 1♂ ( USNM), Vietnam, Hanoi , L. Medvedev @ Dang Dap , fields, gardens, 8.V.1975 ; 3♀ ( BMNH), Vietnam, Tonkin: Thai–Nien. Bank of Fleuve Rouge. , 1924, H. Stevens ; 2♂ 3♀ ( USNM), South India, Western Ghats Karnataka Prov , env. Mudigere , 14.XI. 2003, 904m, river, leg. Konstantinov, Prathapan, Saluk ; 6 ( BMNH), H. L. Andrewes, India, Nilgiri Hills Andrewes, Bequest, B. M. 1922–221, Chaetocnema modesta Gressitt & Kimoto, Det. Yongying Ruan, 2016 .

Remarks: Chaetocnema modesta is rather unique in Oriental Chaetocnema species by the small size of pronotum and the strongly narrowed and constricted pronotal base.

Figures

FIGURE 100. Chaetocnema modesta. A Male habitus. B Pronotum. C Head, frontal view. D Aedeagus, ventral view. E Aedeagus, lateral view. F Apex of aedeagus, ventral view. G Vaginal palpi. H Spermatheca. I Pygidium (tergite 7) of female.