Intermaria, Diana Delicado, ladimir Pešić & Peter Glöer, 2016

Diana Delicado, ladimir Pešić & Peter Glöer, 2016, Unraveling a new lineage of Hydrobiidae genera (Caenogastropoda: Truncatelloidea) from the Ponto-Caspian region, European Journal of Taxonomy 208, pp. 1-29 : 7

Genus Intermaria gen. nov.

urn:lsid:zoobank.org:act:1CDF6CD1-5D3C-447F-BAAE-7197FBEB 2614

Diagnosis

Shell ovate-conic, 3 to 4.5 mm high; large and convex body whorl; rest of whorls small and slightly convex; aperture pyriform, angled on top and often fused to the body whorl. Operculum corneous, yellowish, thin, pliable, ellipsoidal, paucispiral with submarginal nucleus. Two pairs of basal cusps on radular central tooth. Ctenidium occupying nearly the entire length of pallial cavity bearing welldeveloped gill filaments. Osphradium opposite approximate middle of ctenidium. Bursa copulatrix lying against the middle section of the albumen gland; pigmented renal oviduct; one elongate seminal receptacle. Prostate gland bean-shaped, about twice as long as wide; exit of the pallial vas deferens from the posterior-most section of the prostate gland and seminal vesicle entering the prostate gland in its middle section; penis simple, gradually tapering, with its distal end tapered and often with a small distal lobe on the inner edge. Nervous system with black pigmentation typically elongated.

Etymology

From Latin inter - (between) - maria (seas), referring to the occurrence of the genus in the continental area between the Mediterranean and the Caspian seas.

Type species

Pseudamnicola zagrosensis Glöer & Pešić, 2009 .

Remarks

Intermaria species, like those of Pseudamnicola , have an ovate-conic simple shell, one elongated seminal receptacle and a pigmented renal oviduct and nervous system. However, Intermaria differs in having smaller shell dimensions (e.g., when compared with the 5 mm height of P. granjaensis Glöer & Zettler, 2007 see Delicado et al. 2014), a more conic-shaped shell, two basal cusps on the central radular tooth (one in Pseudamnicola ), a shorter prostate gland, and a gradually tapering penis with a small distal lobe on the inner edge (penis is triangular with many surface folds and has a blunt end in Pseudamnicola see Delicado et al. 2015).