Promalactis subquadrata Wang, 2019

Wang, Shuxia & Jia, Yanyan, 2019, Taxonomic study of the genus Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) IV. The sakaiella species group, with descriptions of fifteen new species, Zootaxa 4563 (3), pp. 491-515 : 496-497

Promalactis subquadrata Wang , sp. nov.

( Figs 7 , 20 )

Type material. CHINA , Guizhou Province: Holotype ♂, Dongdai, Shuizu Town (25.36°N, 107.96°E), Limingguan , Libo County, 720 m, 19.VII.2015, coll. Meiqing Yang & Jia’en Li, slide No. JYY17553.

Diagnosis. This new species can be distinguished from other known species of the sakaiella species group in the male genitalia by the costa having a sub-quadrate subapical process that bears a tuft of spines, a small semicircular uncus, and a large linguitate gnathos that is approximately three times the length of the uncus.

Description. Adult ( Fig. 7 ) wingspan 8.5 mm–9.0 mm. Head with vertex creamy white, frons greyish white, occiput dark ferruginous. Labial palpus with first and second segments orange yellow on inner surface, ocherous brown on outer surface; third segment from ocherous brown becoming blackish brown except white at base, slightly longer than second segment. Antenna with scape creamy white; flagellum blackish brown on ventral surface, basal four flagellomeres white, remaining flagellomeres blackish brown annulated with white on dorsal surface. Thorax black; tegula dark ferruginous. Forewing pale ocherous yellow; basal fascia dark ferruginous mixed with deep grey scales, with two parallel white lines: subbasal line edged with black scales, another line along outer margin, edged with black scales along its inner side; costal margin with deep silvery grey scales diffused from middle to before distal 1/3, forming an ill-defined silvery grey spot; cell with a small black spot at middle on outer margin; distal patch deep grey mixed with black, diffused and narrowed along termen to tornus; cilia orange yellow except blackish grey along distal part of costal margin and on tornus. Hindwing and cilia grey. Fore- and midlegs yellowish white on dorsal side, tibiae and tarsi black on ventral side, mid tibia with white scale tuft at apex, tarsi white at apex of each tarsomere; hindleg brown dorsally, yellowish white ventrally, tibia with whitish yellow hairs, tarsus marked with white at apex of each tarsomere.

Male genitalia ( Fig. 20 ). Uncus semicircular, very short, about 1/3 length of gnathos. Gnathos weakly sclerotized, broad linguitate, rounded apically. Tegumen divided from posterior 1/4, straight laterally, narrowed anteriorly. Valva sub-rectangular, widest at base, subparallel medially, straight and setose apically; costa with a large sub-quadrate process between basal 3/5 and subapex, bearing a tuft of spines; sacculus broad, narrowed and setose distally. Juxta with lateral lobes extending to near posterior margin of tegumen, curved at middle, rounded at apex; basal lobe thumb shaped, reaching basal 2/5 length of saccus. Saccus slightly longer than uncus, subtriangular, distinctly narrowed to a rounded apex distally. Aedeagus strong, straight, slightly longer than vlava; cornutus spine-like, less than 1/6 length of aedeagus, placed distally.

Female unknown.

Distribution. China (Guizhou).

Etymology. This epithet is from the Latin sub - and quadratus, referring to the sub-quadrate process of the costa.

Figures

FIGURES 5–10. Adults of Promalactis spp. 5. P. campanulata sp. nov., holotype, ♂; 6. P. tibetica sp. nov., holotype, ♂; 7. P. subquadrata sp. nov., holotype, ♂; 8. P. auremacularis sp. nov., paratype, ♂; 9. P. curvitaeniata sp. nov., paratype, ♀; 10. P. lvovskyi sp. nov., holotype, ♂. Scales =1.0 mm

FIGURES 17–20. Male genitalia of Promalactis spp. 17. P. apicidigitata sp. nov., holotype, slide No. JYY17558; 18. P. campanulata sp. nov., holotype, slide No. DZH11040; 19. P. tibetica sp. nov., holotype, slide No. JYY17326; 20. P. subquadrata sp. nov., holotype, slide No. JYY17553. Scales =0.2 mm.