Makarovana Koçak & Kemal, 2008

Bartsch, Ilse, 2015, The genital area of Halacaridae (Acari), life stages and development of morphological characters and implication on the classification, Zootaxa 3919 (2), pp. 201-259 : 223

Makarovana Koçak & Kemal, 2008

(Figs 62 and 63)

Type species. Acanthopalpus hirsutus Makarova, 1978 .

Female. Genital and anal plates fused. GA with three pairs of pgs, each genital sclerite with two sgs ( Bartsch 1978: fig. 26). Three small pairs of internal gac, two situated in middle part of GO, one pair in posterior part (Fig. 62). Ovipositor at rest hardly extending beyond GO. Basis of ovipositor with pair of spines, apex with ten moderately sclerotized spines (Fig. 63), these four anterior and six posterior spines ending with bifid tip. In ovipositor at rest pair of basal spines and two anterior pairs of apical spines visible through genital sclerites, the former pair near anterior margin of GO, the latter near posterior margin. AE without epimeral pores.

Male. Not known.

Juveniles. With two nymphal and one larval stage. Genital and anal plates fused in both proto- and deutonymphs. Deutonymph with two pairs of pgs, two pairs of sgs, and two pairs of internal gac ( Bartsch 1978: fig. 33). GA of protonymph with one pair of internal gac ( Bartsch 1978: fig. 34); pgs and sgs lacking. AE with remnants of epimeral pores. Larval AE with pair of small epimeral pores.

Remarks. Two species have been described. The records are from cold-temperate to Arctic waters of the northern Atlantic and Pacific Ocean ( Bartsch 2009a).