Copobaeninae Abdullah, 1969

Telnov, Dmitry, 2024, On the taxonomy of some South American and Australo-Pacific Anthicidae Latreille, 1819 (Coleoptera) with new subfamily and tribe-rank synonymies, Zootaxa 5501 (3), pp. 401-424 : 404

Copobaeninae Abdullah, 1969

Abdullah (1969: 334).

= Lemodinae Matthews, 1987 syn. nov.

Matthews (1987: 40).

Abdullah (1969) erected a new anthicid subfamily Copobaeninae for the South American genus Copobaenus Fairmaire & Germain, 1863 based on (sequence as in Abdullah (1969), supplemented by me) entire (non-emarginate) compound eyes, wide cranial neck, procoxal cavity closed internally open externally behind, narrowly separated metacoxa (‘subcontiguous’ as of Abdullah (1969)) with internal carina reduced ‘to a narrow-based apophysis’, simple, basally subdentate pretarsal claws, metendosternite with long stalk which is as long as arms, well-developed laminae and anterior tendons present on arms above their junction point with laminae, metathoracic wing with ‘radial cell closed, anal cell absent’, aedeagus with parameres fused, ovipositor with ‘stylus borne on second segment of coxite’. Additionally, in his key to Anthicidae subfamilies in a couplet with Copobaeninae , Abdullah (1969: 330) specifically mentioned ‘ovipositor with 2-segmented coxites’.

Morphology and taxonomy of the Lemodinae was recently treated in detail by Telnov & Degiovanni (2021: 104– 107) and is not repeated here.Adult Copobaenus nobilis Fairmaire & Germain, 1863 , type species of Copobaeninae , is comprised of the same set of morphological features as described for Lemodinae by Telnov & Degiovanni (2021). So, Lemodinae is found to be identical with Copobaeninae in the present study and a new subfamily-rank synonym is introduced.

Additional features of the Copobaeninae supplementary to the list in Telnov & Degiovanni (2021) are given below (the numeration follows and supplements the former authors):

i.xiii.i. Maxilla—galea and lacinia separated, galea well-developed, apically more or less widened and rounded, densely setose;

iv.ii. Mesoventrite, mesoventral intercoxal process—projecting along almost 2/3 of mesocoxal length, apically slightly bilobate (to lock with vertical ridge of metaventrite) or acute;

iv.ii.i. Mesoventrite—mesothoracic glands present, ducts connect to a single pore at the anterior margin of mesoventrite;

vi.vi.i. Metathoracic wing—R cell basally open, a short remnant of RP3+4 present, almost no RP base and no wedge cell in the medial field ( Lawrence et al. 2022);

viii.iii. Abdomen, morphological sternite IX (with spiculum gastrale)—Y-shaped (misinterpreted I-shaped by Telnov & Degiovanni (2021)), with long or short stem;

x.i. Male genital organs—phallobase not present or short and/or fused with tegmen, penis longer than tegmen, extending anteriorly beyond of it.

x.ii. Ovipositor—styli small, borne on incompletely bi-segmented coxites.