Stylokyrtus, Herman, 2010

Herman, Lee, 2010, Generic Revision Of The Procirrina (Coleoptera: Staphylinidae: Paederinae: Pinophilini), Bulletin of the American Museum of Natural History 2010 (347), pp. 1-78 : 66-67

Stylokyrtus , new genus Figure 9

Type species: Oedodactylus errans Sharp, 1876: 337 , designated here.

DIAGNOSIS: Stylokyrtus has four swollen protarsomeres, segment III has a paratergal carina and lacks paratergites, and tergum and sternum III are fused, but see Discussion for this genus. The preceding characters will separate Stylokyrtus from Oedodactylus , which has three inflated protarsomeres, paratergites on segment III, and tergum and sternum III separated. Tergum and sternum VII are separated in Stylokyrtus , which will distinguish the genus from Oedichirus and Palaminus in which tergum and sternum VII are partially or completely fused. The long lateroapical processes of tergum IX of Stylokyrtus will separate it from Procirrus , Neoprocirrus , and Paraprocirrus , which all have short lateroapical processes. Maxillary palpomere 4 is broad and obliquely truncate in Stylokyrtus , but symmetrically to asymmetrically fusiform in Procirrus and Paraprocirrus and fusiform to securiform in Neoprocirrus . The presence of setae on the edge of the posterior margin of the elytra will separate Stylokyrtus from Procirrus , Oedichirus , Palaminus , and Paraprocirrus .

DESCRIPTION: Head (fig. 9) not pedunculate, wider than long; frontoclypeus without subapical, interantennal ridge; postocular lateral margin strongly converging to neck; postocular lateral margin short; basal angles absent; basal margin of head indistinguishable from lateral margin. Neck across nuchal constriction about seven tenths as wide as greatest postocular width of head; [**nuchal groove] 4; [**nuchal ridge]. Eye length longer than postocular length of head. Dorsal surface with deep simple punctation. Ventral surface without postocular groove. Gular sutures separated. [**Gular pubescence and punctation]. Antennomere 11 about as long as 10; apex without spiniform pencil of setae. Man-

4 This structure and other similarly cited structures were unavailable for study.

dibles with apically bifid denticle; [**prostheca]. Maxillary palpus with fourth palpomere longer than third, securiform, compressed. [**Labium, glossae]. [**Hypopharynx]. Labrum without denticles on anterior margin, but with poorly developed, broad lobe adjacent to emargination. [**Epipharynx].

Prothorax (fig. 9) trapezoidal, longer than wide; widest near anterior margin then broadly rounded and moderately strongly convergent to rounded basal margin. Pronotum with simple, dense, moderately coarse punctation; punctation uniform, but for impunctate midlongitudinal strip. Pronotal marginal ridge present. Notosternal suture poorly developed, evident as weak ridge in broad depression; suture and marginal ridge separated. Hypomeron polished and densely punctate. Postprocoxal lobe long and punctate; transverse carina absent; apex of lobes moderately widely separated from one another. Probasisternum with median carina; surface impunctate. Mesospiracular peritreme with anteromedial margin fused to furcasternum. Elytra slightly shorter than pronotum; humeral angle present; posterior edge with row of setae; subapical region without long, thicker, posteromediad direct- ed seta near lateroapical corner. [**Scutellum, setae covered]. Mesosternum without median carina. [**Mesocoxal acetabulum]. [**Intersternal suture].

Profemur with carina near middle of anteroventral surface. Protibia with three combs near middle; tibia without depression on ctenidial surface; apical portion neither constricted or enlarged. Protarsomeres 1–4 inflated; [**ventral setae]; base of tarsomere 1 not surrounded by cupulate protibial apex; [**symmetry of insertion of tarsomere 5 on tarsomere 4]; [**pubescence on ventral surface of tarsomere 5]. Mesotarsomere 1 longer than second and third combined. Metatarsomere 1 about as long as tarsomeres 2–4.

Abdominal segments without imbricate macrosculpturing. Segment III without paratergites; paratergal carina present basally and laterad of spiracle; tergum and sternum III fused. Segment VII with tergum and sternum separated. [**Segments IV to VI, ‘‘windows’’]. [**Sternum I]. [**Sterna IV–VI, glandular lobes]. Tergum IX [**base]; lateroapical process long, strongly curved ven- trally, and extending well beyond apex of posterior margin of tergum X; [**base of lateroapical process]. [**Tergum X covered].

[**Female genital sclerites].

[**Aedeagus].

ETYMOLOGY: Stylokyrtus , from the Greek for ‘‘pillar’’ or ‘‘column’’ (stylos) and ‘‘curved’’ (kyrtos), refers to the strongly curved lateroapical processes of abdominal segment IX. The generic name is masculine.

DISTRIBUTION AND HABITAT: The genus is found in Brazil, but nothing is known about the habitat of the only species.

DISCUSSION: Stylokyrtus errans was originally described in Oedodactylus , but is removed from that genus because, unlike species in the latter, S. errans has four inflated protarsomeres, abdominal segment III lacks paratergites and has a basal paratergal carina, and tergum and sternum III are fused. The species of Oedodactylus have three inflated protarsomeres and abdominal segment III has a pair of paratergites and the tergum and sternum are separated. The male holotype of the type species was not dissected.

Note, however, that the paratergal carina of S. errans is strongly and coarsely developed and similar to the paratergite of species of Oedodactylus . I cannot see the sutures that delimit a paratergite for segment III and without dissection I am uncertain whether this structure is a carina or a paratergite or if the tergum and sternum are separated or fused. As the species is known only by the holotype, the needed dissection was impossible. Nonetheless, S. errans has four inflated protarsomeres and species of Oedodactylus have only three.

SPECIES INCLUDED AND

MATERIAL EXAMINED

errans (Sharp) – H ( BMNH).......... Brazil [Newcombination,transferredfrom Oedodactylus ]

UNDETERMINED SPECIMENS: None.

DISSECTIONS: None.